- Author: Kathy Keatley Garvey
If you've ever been to Bodega Head in Sonoma County, you may have marveled at the waves crashing and the whales surfacing.
But have you ever seen the digger bees, Anthophora bomboides stanfordiana, aka bumble bee mimics, that nest in the sandstone cliffs?
They're there. They're foraging on flowers, excavating their nests, and rearing young.
These digger bees are not easy to photograph. On our May 9th trip, we got lucky: our 200mm macro lens picked up a digger bee warming its flight muscles.
"The species name indicates that it is a bumble bee mimic," the late Robbin Thorp (1933-2019), a global authority on bumble bees and a UC Davis distinguished emeritus professor of entomology. "These bees need a source of fresh water nearby. Females suck up water, regurgitate it on the sandstone bank surface, then dig away at the soft mud. They use some of the mud to build entrance turrets, presumably to help them locate their nests within the aggregation of nests."
"The female," Thorp said, "sucks up fresh water from nearby, stores it in her crop (like honey bees store nectar) for transport to the nest. She regurgitates it on the sandstone, and excavates the moistened soil. She carries out the mud and makes the entrance turret with it."
Thorp, a 30-year member of the Department of Entomology and Nematology, studied these bees. (See his presentation at the Proceedings of the Symposium on Biodiversity of Northwestern California, Santa Rosa, delivered in October 1991.)
Today, community ecologist and associate professor Rachel Vannette from the same department, is among scientists engaged in the research of these fascinating bees.





- Author: Kathy Keatley Garvey
The open-to-the-public competition involved searching for the first-of-the-year bumble bee in the two-county area of Yolo or Solano; photographing it; and emailing the image to the Bohart Museum at bmuseum@ucdavis.edu.
The first bumble bee to emerge in this area is usually the black-tailed bumble bee, Bombus melanopygus, but another bumble bee, the yellow-faced bumble bee, Bombus vosnesenskii,is out early as well.
Both are considered "spring bees" because that is when their population is the highest, according to Thorp. Then their numbers "tail" the rest of the year.
Lynn Kimsey, director of the Bohart and a UC Davis distinguished professor, Department of Entomology and Nematology, announced Saturday that "we have the winners."
Not one winner, but two. No one species, but two.
Coincidentally, they each took their photos at exactly 2:30 p.m., Jan. 1 in the 100-acre UC Davis Arboretum and Public Garden as the bees foraged on manzanita.
And fittingly, they both knew and worked with Robbin Thorp (1933-2019), a global authority on bees and a UC Davis distinguished emeritus professor of entomology.
The event also marked the second consecutive year that a member of the Williams lab won. Last year postdoctoral researcher Charlie Casey Nicholson of the Williams lab and the lab of Elina Lastro Niño, claimed the prize by photographing a B. melanopygus at 3:10 p.m., Jan. 14 in a manzanita patch in the Arboretum.
As the 2022 winners, Page and Zagory will each receive a coffee cup designed with the endangered Franklin's bumble bee, a bee that Thorp closely monitored in its small range at the California-Oregon border. The cup features the image of the bee specimen, photographed by Bohart scientist Brennen Dyer, and designed by UC Davis doctoral alumnus Fran Keller, professor at Folsom Lake College.
The television program, Good Day Sacramento, featured the contest on Jan. 3. (See it here.)
Worldwide Collection. The Bohart Museum houses a worldwide collection of 8 million insect specimens, including 112 species of bumble bees, Kimsey said. Thorp spend much of his time at the Bohart where he identified bees and helped colleagues with their research.
Thorp, a 30-year member of the UC Davis faculty, retired in 1994 but continued his work on bees until his death at age 85 at his home in Davis. Known as a tireless advocate of pollinator species protection and conservation, he co-authored two books in 2014: Bumble Bees of North America: An Identification Guide (Princeton University) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday).
Said Zagory: “I indeed knew Robbin Thorp, one of the most generous and kind people I have ever met. Dr. Thorp invited me to do a page for their book (California Bees And Blooms: A Guide for Gardeners and Naturalists) about the UC Davis Arboretum All-Stars (pages 230 -232) and he edited a publication we created at the UC Davis Arboretum called Ten Bees and Ten Plants they Love that can be downloaded from the website at https://arboretum.ucdavis.edu/pollinator-gardening.
Page is a 2018 alumnus of The Bee Course, which Thorp co-taught from 2002-2018. The nine-day intensive workshop, geared for conservation biologists and pollination ecologists and considered the world's premiere native bee biology and taxonomic course, takes place annually in Portal, Ariz. at the Southwestern Research Station, part of the American Museum of Natural History, N.Y. (The 2021 winner--Charlie Casey Nicholson--is a 2015 alumnus of The Bee Course.)
Page said she was “also lucky enough to participate in a "Bumble Bee Blitz" organized by Thorp and the U. S. Fish and Wildlife Service in July 2016 on Mt. Ashland, where we searched for Bombus franklini and Bombus occidentalis-- two very rare West Coast bee species. We unfortunately did not find Bombus franklini, which is now recognized as an endangered species under the Endangered Species Act.”
Brilliant Scientist. “Robbin was a brilliant scientist and a dedicated advocate for bumble bee conservation,” Page said. “His death was a great loss and I wish more of my career could have overlapped with his time in Davis.”
As a doctoral candidate in entomology, Page researches and investigates “whether European honey bees compete with native bees for floral resources and how we can use well-planned floral enhancements to mitigate negative effects of competition."
How rare is it find Bombus vosnesenskii on Jan. 1?
In an email today, UC Davis doctoral alumnus and Thorp protégé Kim Chacon, said she has seen B. vosnesenskii near San Luis Obispo since Dec 26. She is a lecturer at California Polytechnic University (Cal Poly). "I think it is a very opportunistic species. In my research, that was one of the first emergers, the 6th genus actually. Robbin Thorp had an interesting theory about bumble bees dealing with a virus or other illness which was shifting the dominant species to vosnesenskii. I'm so happy both Ellen and Maureen won-- they are both awesome!"
"My congratulations, too," said bumble bee enthusiast/photographer Allan Jones of Davis, a friend of Thorp's. "I did not even go out considering the chilly weather. I expected it to be on the second day when we got up into the sixties, and with the ground so damp and cold, too. Hats off, brrrr."
Both bumble bee species have also been sighted and photographed in recent years on Dec. 31 and Jan. 1 in Benicia, Solano County.



- Author: Kathy Keatley Garvey
If you're thinking about taking a walk in Yolo or Solano counties to celebrate the new year--or just to get some exercise in keeping with your New Year's Resolution (you did make one, didn't you?)--bring your camera.
You might find and photograph the first bumble bee of the year in a contest sponsored by the Bohart Museum of Entomology and memorializing bumble bee expert Robbin Thorp (1933-2019), UC Davis distinguished emeritus professor of entomology.
Participants are to capture an image of a bumble bee in the wild in either of the two counties and email the image to bmuseum@ucdavis.edu, with the details of time, date and place. The image must be recognizable as a bumble bee, said contest coordinator Lynn Kimsey, director of the Bohart Museum and a UC Davis distinguished professor of entomology.
The first bumble bee to emerge in this area is the black-tailed bumble bee, Bombus melanopygus, according to Thorp, who co-authored Bumble Bees of North America: An Identification Guide (Princeton University) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday).
Thorp launched the first unofficial bumble-bee-of-the-year contest several years ago with a small group of bumble bee enthusiasts/photographers: Allan Jones and Gary Zamzow of Yolo County, and yours truly of Solano County. Kim Chacon, then a UC Davis doctoral candidate and a Thorp protégé, joined us later. She is a 2019 alumnus of The Bee Course (co-taught by Thorp from 2002-2018). The nine-day intensive workshop, geared for conservation biologists and pollination ecologists and considered the world's premiere native bee biology and taxonomic course, takes place annually in Portal, Ariz. at the Southwestern Research Station, part of the American Museum of Natural History, N.Y.
Nicholson photographed B. melanopygus, in a manzanita patch at 3:10 p.m., Thursday, Jan. 14, 2021 in the UC Davis Arboretum and Public Garden to claim the honor. He is a researcher in the UC Davis Department of Entomology and Nematology labs of Professor Neal Williams, a pollination ecologist, and Extension apiculturist Elina Lastro Niño.
His prize? A coffee cup designed with the endangered bumble bee that Thorp closely monitored—Franklin's bumble bee, Bombus franklini, known to exist in a small area by the California-Oregon border. UC Davis doctoral alumnus Fran Keller, a professor at Folsom Lake College and a Bohart Museum scientist, designed the cup. Bohart scientist Brennen Dyer photographed the specimen from the Bohart collection.
The 2022 winner also will receive that prized coffee cup.
So, take a walk. Invite your camera. Keep that New Year's Resolution.
Game on!

- Author: Kathy Keatley Garvey
What's it all about? The Bohart Museum of Entomology is launching its second annual Robbin Thorp Memorial Bumble-Bee-of-the-Year Contest to see who can find and photograph the first bumble bee of 2022 in Yolo or Solano counties.
Participants are to capture an image of a bumble bee in the wild in either of the two counties and email the image to bmuseum@ucdavis.edu, with the details of time, date and place. The image must be recognizable as a bumble bee, said contest coordinator Lynn Kimsey, director of the Bohart Museum and a UC Davis distinguished professor of entomology.
The winner will receive a coffee cup designed with the endangered bumble bee that the late Robbin Thorp closely monitored—Franklin's bumble bee, Bombus franklini, known to exist in a small area by the California-Oregon border. UC Davis doctoral alumnus Fran Keller, a professor at Folsom Lake College and a Bohart Museum scientist, designed the cup. Bohart scientist Brennen Dyer photographed the specimen.
Thorp, a global authority on bees and a distinguished emeritus professor of entomology at UC Davis, always looked forward to finding or seeing the first bumble bee of the year in the area.
The native black-tailed bumble bee, Bombus melanopygus, is the first bumble bee to emerge in the area, according to Thorp. It forages on manzanitas, wild lilacs, wild buckwheats, lupines, penstemons, clovers, and sages, among others.
Thorp served on the UC Davis entomology faculty for 30 years, from 1964 to 1994. Although he achieved emeritus status in 1994, he continued to engage in research, teaching and public service until a few weeks before his death on June 7, 2019 at age 85 at his home in Davis.
Nicholson, a researcher in the UC Davis Department of Entomology and Nematology labs of Professor Neal Williams, a pollination ecologist, and Extension apiculturist Elina Lastro Niño, is a 2015 alumnus of The Bee Course, where Thorp taught from 2002 through 2018. The nine-day intensive workshop, geared for conservation biologists and pollination ecologists and considered the world's premiere native bee biology and taxonomic course, takes place annually in Portal, Ariz., at the Southwestern Research Station, part of the American Museum of Natural History, N.Y.
Kimsey praised Thorp for his expertise, generosity and kindness. Kimsey, who first met Thorp when she was a graduate student at UC Davis, said that although he wasn't her major professor, “my project was on bees and he was incredibly helpful and supportive. His enthusiasm about pollinators and bees in particular actually grew after he retired, and he continued helping students and researchers and was the backbone of so much research. His support and kindness was matched by his undemanding assistance and expertise.”
In 2014, Thorp co-authored two books, Bumble Bees of North America: An Identification Guide (Princeton University) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday).
Thorp, the last known person to see Franklin's bumble bee in its native habitat, spotted it in 2006 near Mt. Ashland. The bee inhabits--or did--a 13,300-square-mile area within the five-county area of Siskiyou and Trinity in California; and Jackson, Douglas and Josephine in Oregon.
Thorp sighted 94 Franklin's bumble bees in that area in 1998, but by 2003, the tally had dropped to three. Thorp saw none in 2004 and 2005; one in 2006; and none since. Thorp's determined hunt for the bumble bee resulted in the CNN publication of "The Old Man and the Bee," a spin-off of Hemingway's "The Old Man and the Sea."

- Author: Kathy Keatley Garvey
Back in the 2009, a team of University of California scientists, led by Gordon Frankie of UC Berkeley and Robbin Thorp (1933-2019) of UC Davis, wrote this incredible article, "Native Bees are a Rich Natural Resource in Urban California Gardens," published in California Agriculture (UC Agriculture and Natural Resources).
It's an excellent resource.
The abstract:
"Evidence is mounting that pollinators of crop and wildland plants are declining worldwide. Our research group at UC Berkeley and UC Davis conducted a 3-year survey of bee pollinators in seven cities from Northern California to Southern California. Results indicate that many types of urban residential gardens provide floral and nesting resources for the reproduction and survival of bees, especially a diversity of native bees. Habitat gardening for bees, using targeted ornamental plants, can predictably increase bee diversity and abundance, and provide clear pollination benefits."
Jennifer Hernandez, Barbara Ertter, Jaime Pawelek, Sara Witt, Mary Schindler, Rollin Coville, and Victoria Wojcik. Coville provided the images.
"Outdoor urban areas worldwide are known to support a rich diversity of insect life (Frankie and Ehler 1978)," they wrote. "Some insects are undesirable and characterized as pests, such as aphids, snails, earwigs and borers; urban residents are most aware of these. Other urban insects are considered beneficial or aesthetically pleasing, such as ladybird beetles and butterflies; this category includes a rich variety of insects whose roles in gardens go largely unnoticed and are thus underappreciated (Grissell 2001; Tallamy 2009). They regularly visit flowers and pollinate them, an important ecological service."
"We report the results of a 2005-to-2007 survey of bees and their associations with a wide variety of ornamental plant species in seven urban areas, from Northern California to Southern California. While non-native honey bees (Apis mellifera) are common in many gardens, numerous California native bee species also visit urban ornamental flowers. Of about 4,000 bee species known in the entire United States, about 1,600 have been recorded in California."
That deserves to be repeated. Of the 4000 bee species known in the United States, about 1600 reside in California. See some of them listed on the chart above.
Notice it includes the non-native bee, the honey bee, Apis mellfera. It is an "introduced bee," although numerous non-scientists think it's a native. European colonists introduced it to America (in what is now Virginia) in 1622.
The phrase, "Save the (Honey) Bees," became prominent during the colony collapse disorder. But the bees that need saving now are the natives.
If you want to know more about the native bees in California, this book should be in your library: California Bees and Blooms: A Guide for Gardeners and Naturalists by UC-affiliated scientists
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