- Author: Kathy Keatley Garvey
When the UC Davis emeritus professor of entomology, a global authority on bumble bees, died June 7, 2019 at age 85, scientists found a way to memorialize him and what he loved.
The Bohart Museum of Entomology--where Thorp spend much of his time identifying bees, helping scientists, and encouraging guests at open houses to learn about the wonderful world of bees--decided to memorialize him with an annual Robbin Thorp Memorial First-Bumble-Bee-of-the-Year Contest. The first person to photograph a bumble bee in 2021 in the two-county area of Yolo and Solano would win.
And it is only fitting that Charlie Casey Nicholson, who studied bees with Thorp, won. He photographed a black-tailed bumble bee, Bombus melanopygus, in a manzanita patch at 3:10 p.m., Thursday, Jan. 14 in the UC Davis Arboretum and Public Garden to claim the honor. The site is near Old Davis Road.
Due to inclement weather, bumble bees are not easy to find this time of year. Neither are they easy to photograph.
In fact, Nicholson noted this was his seventh observation field trip to look for the first bumble bee of the year. He had searched six previous times (three 10-minute observations on the manzanita on each of two other days, Jan. 6 and 7).
As the winner, Nicholson will receive a special Bohart bumble bee coffee cup and a face mask, said contest coordinator Lynn Kimsey, director of the Bohart Museum and a UC Davis professor of entomology.
"It is truly an honor to win the contest," said Nicholson. "I was a student of Robbin's during the 17th annual Bee Course in Portal, Ariz. I will never forget him wielding his canopy net."
"The first night (8/17/2015) he gave the opening seminar--a whirlwind tour of what makes a bee. It was so exciting to be at this research station surrounded by people whose names you've read all the time.”
“Robbin helped me learn to pay close attention to the arolia of Anthidiini. As we moved into identifying bees, Robbin was a great teacher as we worked through the dichotomous keys in The Bee Genera of North and Central America: Hymenoptera Apoidea. He always had some morphological signpost that wouldn't give away the 'answer' but would certainly guide you in the right direction."
Charlie holds a bachelor of arts degree in biology (evolution, ecology and behavior), 2010, cum laude, from Skidmore College, Saratoga Springs, New York. He received his doctorate in natural resources in 2018 from the University of Vermont, where he was a Gund Institute for Environment graduate fellow. In his dissertation, he examined how landscape and farm management affect the multiple benefits provided by wild bees.
Nicholson joined UC Davis as a postdoctoral scholar in the spring of 2019, and receives funding support from the USDA Invasive Species and Pollinator Health Unit. He recently co-authored a paper, “Natural Hazard Threats to Pollinators and Pollination,” published in the journal Global Change Biology, that analyzed 117 published research papers on natural hazards that threaten pollinators and pollination.
His other interests include multiple dimensions of biodiversity, conservation planning, agricultural management, ecosystem services, and community and landscape ecology.
Robbin Thorp
Thorp, a member of the UC Davis entomology faculty for 30 years, from 1964-1994, died June 7, 2019 at his Davis home at age 85. A tireless advocate of pollinator species protection and conservation, Thorp was known for his expertise, dedication and passion in protecting native pollinators, especially bumble bees, and for his teaching, research and public service. He was an authority on pollination ecology, ecology and systematics of honey bees, bumble bees, vernal pool bees, conservation of bees, native bees and crop pollination, and bees of urban gardens and agricultural landscapes.
He achieved emeritus status in 1994 but continued to engage in research, teaching and public service until a few weeks before his death. In 2014, he co-authored two books Bumble Bees of North America: An Identification Guide (Princeton University, 2014) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday, 2014).
Every summer from 2002 to 2018, Thorp volunteered his time and expertise to be one of the instructors in The Bee Course. In a 2013 interview with the UC Davis Department of Entomology and Nematology, Thorp said he loved teaching at The Bee Course and praised the co-instructors and students. "Ron McGinley who got his undergraduate degree at UC Davis does most of the initial student contact and scheduling for the course. Steve Buchmann, who got his PhD at UC Davis in 1978, is one of the instructors. There are usually about eight instructors and 22 participants for the course. Most of the time is spent in the lab identifying bees to genus. At least three days are spent in the field so students can see various bees doing their thing, collect them and bring them back to the lab to identify them. It is a great experience for students to interact with instructors and especially with their peers from around the world. Instructors all donate their time to teach in the course, but benefit from the chance to get together with colleagues and a new cohort of interesting students each year. Every class is different (that is, it takes on its own personality) and each student brings something new and different to the mix."
Robbin Thorp would have been proud of what happened on Thursday, Jan. 14.



- Author: Kathy Keatley Garvey
Yes, after a l-o-n-g, cold, hard winter, bumble bees are emerging.
At least in Solano County.
At 11:20 a.m. today (Wednesday, Jan. 13), we spotted a yellow-faced bumble bee, Bombus vosnesenskii, foraging on oxalis near downtown Benicia.
What a delight! A January bumble bee and the first one we've seen this year.
So, this is a good time to mention the inaugural Robbin Thorp First-Bumble-Bee-of-the-Year Contest, sponsored by the Bohart Museum of Entomology and coordinated by Bohart director Lynn Kimsey, UC Davis professor of entomology.
Just photograph a clearly identifiable bumble bee in the two-county area of Yolo or Solano, and submit it to bmuseum@ucdavis.edu with the time, date and place. The winner receives bragging rights and a special gift from the Bohart Museum. Plans call for a Bohart coffee mug with a bumble bee image.
The contest memorializes native pollinator specialist and Bohart associate Robbin Thorp (1933-2019), UC Davis distinguished emeritus professor of entomology. Professor Thorp, 85, who died June 7, 2019, was a global authority on bumble bees, and always looked forward to seeing the first bumble bee of the year. He launched an impromptu contest several years ago with a small group of bumble bee enthusiasts/photographers (including yours truly) from Yolo and Solano counties. (Note: I am not a contestant.)
Early Bumble Bees
The black-tailed bumble bee, Bombus melanopygus, is usually the first bumble bee to emerge in this area, Thorp used to say. It forages on manzanitas, wild lilacs, wild buckwheats, lupines, penstemons, clovers, and sages, among others. In early January, it's often seen on manzanita blossoms in the UC Davis Arboretum and Public Garden. B. vosnesenskii forages on such early bloomers as oxalis, jade, mustard and wild radish, and then on a variety of plants throughout the year. Both species are natives.
Thorp, a 30-year member of the UC Davis entomology faculty, from 1964-1994, co-authored two books, Bumble Bees of North America: An Identification Guide (Princeton University, 2014) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday, 2014). He achieved emeritus status in 1994 but continued to engage in research, teaching and public service until a few weeks before his death.
A tireless advocate of pollinator species protection and conservation, Thorp was known for his expertise, dedication and passion in protecting native pollinators, especially bumble bees, and for his teaching, research and public service. He was an authority on pollination ecology, ecology and systematics of honey bees, bumble bees, vernal pool bees, conservation of bees, native bees and crop pollination, and bees of urban gardens and agricultural landscapes.
Ready to join the hunt for the first bumble bee of the year in Yolo and Solano counties? Camera ready? Walking shoes laced? Go!




- Author: Kathy Keatley Garvey
If so, and if you photograph the first one of the year, you might win the Bohart Museum of Entomology's contest.
In memory of native pollinator specialist Robbin Thorp (1933-2019), UC Davis distinguished emeritus professor of entomology, the Bohart Museum is sponsoring the inaugural Robbin Thorp Memorial First-Bumble-Bee-of-the-Year Contest.
Professor Thorp, 85, who died June 7, 2019, was a global authority on bumble bees, and always looked forward to seeing the first bumble bee of the year. He launched an impromptu contest several years ago with a small group of bumble bee enthusiasts/photographers from Yolo and Solano counties.
Now the Bohart Museum, where Thorp spent much of his time identifying bees and helping others, is sponsoring the contest. Participants are to capture an image of a bumble bee in the wild in either Yolo or Solano counties and email the image to bmuseum@ucdavis.edu, with the details of time, date and place. The image must be recognizable as a bumble bee. The winner receives bragging rights and a special gift from the Bohart Museum, said Lynn Kimsey, director of the Bohart and UC Davis professor of entomology. Plans call for a Bohart coffee mug with a bumble bee image.
The first bumble bee to emerge in this area is the black-tailed bumble bee, Bombus melanopygus. Native to western North America and found from California to British Columbia and as far east as Idaho, it forages on manzanitas, wild lilacs, wild buckwheats, lupines, penstemons, clovers, and sages, among others.
Thorp, a 30-year member of the UC Davis entomology faculty, from 1964-1994, co-authored two books, Bumble Bees of North America: An Identification Guide (Princeton University, 2014) and California Bees and Blooms: A Guide for Gardeners and Naturalists (Heyday, 2014). He achieved emeritus status in 1994 but continued to engage in research, teaching and public service until a few weeks before his death.
A tireless advocate of pollinator species protection and conservation, he was known for his expertise, dedication and passion in protecting native pollinators, especially bumble bees, and for his teaching, research and public service. He was an authority on pollination ecology, ecology and systematics of honey bees, bumble bees, vernal pool bees, conservation of bees, native bees and crop pollination, and bees of urban gardens and agricultural landscapes.
Thorp co-taught The Bee Course from 2002 to 2019, an intensive nine-day workshop affiliated with the American Museum of Natural History and held annually at the Southwestern Research Station, Portal, Ariz.
Kimsey, who first met Thorp when she was a graduate student at UC Davis, said that although he wasn't her major professor, “my project was on bees and he was incredibly helpful and supportive. His enthusiasm about pollinators and bees in particular actually grew after he retired, and he continued helping students and researchers and was the backbone of so much research. His support and kindness was matched by his undemanding assistance and expertise. What a terrible loss to his family and to the research and conservation communities."
An authority on the critically imperiled Franklin's bumble bee, Bombus franklini, Thorp began monitoring the bumble bee population in 1998 in its narrow distribution range of southern Oregon and northern California. He had not seen it since 2006 and was instrumental in placing Franklin's bumble bee on the Red List of Threatened Species of the International Union for Conservation of Nature and Natural Resources (IUCN). Long active in the North America IUCN Bumblebee Specialist Group, Thorp served as its regional co-chair, beginning in 2011.
In August of 2016 a documentary crew from CNN, headed by John Sutter, followed Thorp to a meadow where he last saw Franklin's bumble bee. Sutter wrote about Dr. Thorp, then 82, in a piece he titled "The Old Man and the Bee," aspinoff of Ernest Hemingway's "The Old Man and the Sea." (See "Will Franklin's Bumble Bee Ever Be Seen Again?"on YouTube by EarthFixMedia.)
Highly honored by his peers, Thorp was named a fellow of the California Academy of Sciences, San Francisco in 1986; recipient of the Edward A. Dickson Emeriti Professorship of UC Davis in 2010; and recipient of the UC Davis Distinguished Emeritus Award in 2015. Other honors included: member of the UC Davis Bee Team that won the Team Award from the Pacific Branch, Entomological Society of America (PBESA) in 2013. In addition, he was a past president (2010-2011) of the Davis Botanical Society, and former chair (1992-2011) of the Advisory Committee for the Jepson Prairie Reserve, UC Davis/Natural Reserve System.
UC Davis professor and pollination ecologist Neal Williams, who organized a special symposium in Thorp's honor at the 2019 PBESA meeting in San Diego, praised his “tireless efforts in research, advocacy and education” and how he “inspired a new generation of bee researchers.”



- Author: Kathy Keatley Garvey
What was that foraging on a pink iceplant blossom near a path to the ocean? A metallic green sweat bee, Agapostemon texanus, also called an ultra green sweat bee.
We usually don't see A. texanus unless it's spring or summer, but there it was, out of season. Or rather, there "he" was. Males and females are easily distinguishable. The female is solid green, from head to thorax to abdomen, while the green coloration on the male appears on the head and thorax.
We remember pollinator specialist Robbin Thorp (1933-2019) emeritus professor of entomology at UC Davis, talking about them. He delighted in seeing them at his monitoring site, the Department of Entomology and Nematology's Häagen-Dazs Honey Bee Haven on Bee Biology Road, UC Davis campus. The Agapostemon are members of the Halictinae family. They are often called "sweat bees" because they are attracted to human sweat, probably for the salt.
Green sweat bees are among the bees featured in the book, "California Bees and Blooms: A Guide for Gardeners and Naturalists," co-authored by the University of California team of Gordon Frankie, UC Berkeley; Robbin Thorp, UC Davis; and UC Berkeley affiliates Rollin Coville (photographer and entomologist) and Barbara Ertter (plant specialist). Frankie, Thorp, Coville and Ertter (and others) also published "Native Bees Are a Rich Natural Resource in Urban California Gardens" in California Agriculture.



- Author: Kathy Keatley Garvey
Have you ever pulled up a chair in your garden and watched honey bees foraging?
They are so intent on their "bees-ness" that they don't know you're there. It's a great opportunity to photograph them.
Sometimes, if you're lucky, they'll buzz over your head on their way back to their colony, and you'll see:
- The three main body parts: head, thorax and abdomen
- The two pairs of wings
- The three pairs of legs
- The pair of antennae
Such was the case in Vacaville this week when we were watching honey bees forage in our African blue basil, a bee magnet that we plant annually. We first learned of African blue basil, (Ocimum kilimandscharicum × basilicum 'Dark Opal'), through Gordon Frankie, UC Berkeley professor and the late Robbin Thorp, distinguished emeritus professor of entomology at UC Davis. They co-authored the book, California Bees and Blooms: a Guide for Gardeners and Naturalists (Heyday Books) with Rollin Coville and Barbara Ertter, also affiliated with UC Berkeley.
Want to know more about honey bees? Be sure to read the newly published The Art of the Honey Bee; Shaping the Environment from Landscapes to Societies (Oxford University Press) by noted bee geneticist and biologist Robert E. Page Jr., who maintains strong ties to UC Davis and Arizona State University (ASU). Also learn about honey bee anatomy on ASU's web page, "Ask a Biologist."
Page, who holds a doctorate in entomology from UC Davis, is a former chair of the UC Davis Department of Entomology. In 2004, Arizona State University (ASU) recruited him for what would become a series of top-level administrative roles. He advanced from director of the School of Life Sciences to dean of Life Sciences; vice provost and dean of the College of Liberal Arts and Sciences; and university provost. Today he holds the titles of provost emeritus of ASU and Regents professor emeritus, as well as UC Davis department chair emeritus, professor emeritus, and UC Davis distinguished emeritus professor.
Tool Kit
Did you know a bee has a tool kit? Page lists the tool kit in his book, The Art of the Bee: a compass, an odometer and a path integrator.
'As 'central place foragers,' bees fly out from the nest site and explore the surrounding environment in search of food resources," writes Page, renowned for his research on honey bee behavior and population genetics, particularly the evolution of complex social behavior. "They return to the nest with the resources they collect. To do this, they need to be able to navigate out and find their way back. To aid them, they have a toolkit of navigation mechanisms."
One tool in their tool kit is their internal compass that depends on the location of the sun.
"As light from the sun passes through the atmosphere, it becomes polarized," Page writes. "The pattern of polarized light in the sky depends on the angle of the sun relative to where you are looking. Bees have special sensors in their eyes for detecting the polarized light patterns. On cloudy days, they can't see the sky; but they can still locate the sun using ultraviolet light detectors. Ultraviolet light penetrates cloud cover, allowing bees to use the location of the sun as a navigational marker. With heavy clouds, bees can get to and from a resource by relying solely on landmarks that they learn; otherwise, they stay home until the weather changes. However, as the earth turns, the sun is always changing location relative to the horizon, making it an unreliable marker unless you know the time of day, and bees do. They learn the movement of the sun across the sky and reference it to an internal clock. We know they have the clock because we can train them to forage at specific times of day. If you anesthetize a bee, you can stop her clock. When she awakens and takes a foraging trip to a learned foraging station, her flight path will be offset by the amount of time lost. In other words, she will misinterpret the direction based on the current location of the sun by the amount of time she was anesthetized."
"The odometer plus the ability to determine a flight vector (direction and distance) from a given landmark along a resource flight path, using their sun compass and internal clock, give bees the basic tools for navigation," Page writes. "The last tool in the toolkit is a path integrator that combines the compass and odometer information."
It's a fascinating book by Page, whose most salient contributions to science include constructing the first genomic map of the honey bee, which sparked a variety of pioneering contributions not only to insect biology but to genetics at large.
Meanwhile, take the challenge. Pull up a chair in your garden and watch and photograph the bees going about their "bees-ness."



