- Author: Kathy Keatley Garvey
The deadline to RSVP for the Celebration of Life for internationally known honey bee authority, Extension apiculturist Eric Mussen of the UC Davis Department of Entomology and Nematology, is Monday, Aug. 1.
The memorial is set from 4 to 6 p.m., Sunday, Aug. 28 in the Putah Creek Lodge, 685 Putah Creek Lodge Drive. To RSVP, access https://ericmussencelebrationofli.rsvpify.com. For those who cannot attend in person, UC Davis distinguished professor Walter Leal of the Department of Molecular and Cellular Biology, and a former chair of the Department of Entomology, is generously donating his time and talents to do a live Zoom webinar. Registration is underway here at https://bit.ly/3czl5Am; no deadline to register. It also will be on YouTube.
Dr. Mussen, a 38-year California Cooperative Extension apiculturist died Friday, June 3 from liver cancer, after a diagnosis several days before. He was 78. (See https://bit.ly/3ou1O5W)
The tentative program:
- Introductory remarks by administrators, including UC Davis Chancellor Gary May
- Speakers from UC Davis, the bee industry and almond industry
- Memory table
- PowerPoint presentation of images
- Songs by the Davis-based Doo wop group directed by Frank Fox (Dr. Mussen was an active member)
- A honey bee observation hive, compliments of Extension apiculturist Elina Lastro Niño, director of the California Master Beekeeper Program
- Light refreshments, including a bee-themed cake.
The National Honey Board is donating honey bee pins, and Amina Harris, director of the Honey and Pollination Center, will provide honey straws.
Dr. Mussen, known to all as "Eric,” joined the UC Davis entomology department in 1976. Although he retired in 2014, he continued his many activities until a few weeks prior to his death. For nearly four decades, he drew praise as “the honey bee guru,” “the pulse of the bee industry" and "the go-to person" when consumers, scientists, researchers, students, and the news media sought answers about honey bees. Join us to celebrate his life and legacy.
Dr. Mussen worked with everyone interested in honey bees--from 4-H beekeepers to large-scale commercial beekeeping operations. For years, he volunteered his expertise for the National 4-H Beekeeping Essay Competition sponsored by the Foundation for the Preservation of Honey Bees. He chaired the California-level contest and judged the entries.
Family and friends suggest memorial contributions be made to the California State 4-H Beekeeping Program, with a note, "Eric Mussen Memorial Fund." Mary Ciriceillo, director of development for the UC Agriculture and Natural Resources, said checks may be made out to the California 4-H Foundation and mailed to:

- Author: Kathy Keatley Garvey
Jean-Pierre Delplanque, vice provost and dean of Graduate Studies, says the program recognizes "faculty providing outstanding service in advising and mentoring at the program level."
In a letter to Professor Rosenheim, he praised his "excellent service to your graduate program, as well as your positive impact on graduate students and your colleagues. We thank you for your investment in advising and mentoring graduate students and contribution to their success."
Delplanque singled out a few excerpts from the award packet:
- “He has demonstrated his unparalleled dedication to mentoring students, and ability to cultivate an atmosphere of learning, excitement, and critical thinking in the lab, field, and classroom. He consistently makes time to support his students. Regardless of his current teaching and research demands, Jay will provide feedback, schedule meetings, and maintain his open door policy.”
- “Jay is a remarkably skillful and reassuring mentor with a natural generosity of spirit and a broad view of mentorship. He has deep knowledge in a wide range of subject areas, and also has the wisdom to offer students good advice on all stages of the research process. Jay is especially good at putting students at ease and motivating them to persist through the setbacks of research. I have often seen students go to Jay for advice when they are worried and leave feeling better about the road ahead.”
Delplanque noted that "Graduate Studies is committed to showcasing and promoting positive mentoring experiences like yours. Graduate advising and mentoring is vital for guiding students through their degrees and professional development, while also helping ensure their overall success and well-being. Your efforts exemplify outstanding service in mentorship and we hope you will join us in championing the benefits and significance of graduate student mentoring across campus. Congratulations again on your achievement."
Rosenheim, who specializes in insect ecology, integrated pest management, and biological control, and the use of farmer-generated data to enhance pest and crop management ('Ecoinformatics'), holds a bachelor's degree in entomology (1983) from UC Davis and a doctorate in entomology (1987) from UC Berkeley. He joined the UC Davis faculty in 1990. and become a UC Davis distinguished professor in 2018.
Humbled and honored to receive the award, Rosenheim says that "The job of a professor is quite diverse, and quite rewarding in different ways. Teaching in a classroom provides instant gratification, as you see the light of understanding and excitement shining in students' eyes as they explore and grasp new concepts. Research in the laboratory provides instead delayed gratification, where long periods of hard work--sometimes years--may pass before questions are answered and one feels the satisfaction of pushing forward the margins of scientific understanding."
"But, perhaps the most lasting sense of accomplishment comes from mentoring graduate students," Rosenheim says. Building relationships with graduate students, watching them grow in their skills and confidence and, finally, seeing them establish themselves in their careers, provides the kind of reward that is similar in some ways to the happiness that parents derive from their children. And the relationships never end – they are bonds that last a lifetime. I think the key to effective mentorship is to place the student's welfare at the top of one's priority list. So, drafts of papers should be returned promptly with constructive suggestions, and not allowed to languish in a long queue of manuscripts waiting for reviews--more senior colleagues can wait, if someone needs to wait.
"And, I think effective mentorship also means tailoring the kind of assistance provided to the needs and desires of each student. Mentorship is definitely not a one-size-fits-all kind of undertaking. Some students need a lot of encouragement (actually, almost everyone benefits from positive feedback, because research often produces huge servings of critiques), some students need more assistance, especially at the earliest stages of their research, but some students absolutely chafe under too much input, and instead want total independence. That's fine – it's important to adjust to each student's desires and needs. I think good mentorship also means establishing a laboratory culture of openness and collegiality. Everyone should be happy to come to the lab, learn from each other, and contribute to each other's research progress. Mentorship doesn't just come from the lab PI (principal investigator), but from everyone who makes the lab a community."
A native of Yorktown, N.Y, young Jay developed an interest in biology while exploring the vernal pools behind his Hudson River Valley home. As an undergraduate at UC Davis, he initially majored in physics. "On a lark" he enrolled in Professor Harry Kaya's Entomology 100 course in 1981. The professor inspired him, the class enthralled him, and insects captivated him.
Rosenheim's career has not only led to his being elected a Fellow of the Entomological Society of America (ESA) and a Fellow of the American Association for the Advancement of Science, but recipient of teaching awards from the Associated Students of UC Davis and the UC Davis Academic Senate; and the Distinguished Student Mentoring Award from ESA's Pacific Branch.
He co-founded and co-directs the campuswide Research Scholars Program in Insect Biology (RSPIB) with Professors Joanna Chiu and Louie Yang of the UC Davis Department of Entomology and Nematology. The program aims to provide "undergraduates with a closely-mentored research experience in biology," according to the website. "Because insects can be used as model systems to explore virtually any area of biology (population biology; behavior and ecology; biodiversity and evolutionary ecology; agroecology; genetics and molecular biology; biochemistry and physiology; cell biology), faculty in the program can provide research opportunities across the full sweep of biology. The program's goal is to provide academically strong and highly motivated undergraduates with a multi-year research experience that cultivates skills that will prepare them for a career in biological research."

- Author: Kathy Keatley Garvey
The celebration of life will take place from 4 to 6 p.m., Sunday, Aug. 28 in the Putah Creek Lodge, UC Davis campus. The deadline for reservations is Aug. 1. RSVP here: https://ericmussencelebrationofli.rsvpify.com.
“Eric was a giant in the field of apiculture,” said Steve Nadler, professor and chair of the UC Davis Department of Entomology and Nematology. “The impact of his work stretched far beyond California.”
The event will include speakers, a PowerPoint of images showcasing his life, a memory table, songs by the Davis doo wop chorus he belonged to, and light refreshments. The crowd, expected to fill thePutah Creek Lodge, will include family and friends; UC Davis faculty, staff and students; bee and almond industry colleagues; and his musician friends. For those unable to attend, the event will include a webinar (register here) produced by UC Davis distinguished professor Walter Leal of the Department of Molecular and Cellular Biology, a former chair of the Department of Entomology. It also will be on YouTube.
Dr. Mussen, known to all as “Eric,” joined the UC Davis entomology department in 1976. Although he retired in 2014, he continued his many activities until a few weeks prior to his death. For nearly four decades, he drew praise as “the honey bee guru,” “the pulse of the bee industry" and as "the go-to person" when consumers, scientists, researchers, students, and the news media sought answers about honey bees.
Colleagues described Eric as the “premier authority on bees and pollination in California, and one of the top beekeeping authorities nationwide,” “a treasure to the beekeeping industry," and "a walking encyclopedia when it comes to honey bees."
When Eric was nominated (and received) the 2013-14 Distinguished Service Award for Outstanding Extension from the UC Agriculture and Natural Resources (UC ANR), MacArthur Genus Awardee Professor Marla Spivak, Distinguished McKnight University Professor Apiculture/Social Insects at the University of Minnesota, wrote: "Without question, Eric is the No. 1 Extension person dealing with honey bees in the nation, if not the world. Research colleagues, beekeepers and the public are all very lucky to have him.” (See tribute on UC Davis Department of Entomology and Nematology website.)
In his leisure time, Eric sang doo wop for more than a decade with a performance class taught by Frank Fox of Davis. "The class taught was always a performance class and culminated in a concert--the Davis Arts Center Doowop Ensemble was what I called it," Fox said. "Eric sang bass but often did leads as well." Among Eric's favorites: "The Duke of Earl," "Blue Moon, "All I Have to Do is Dream" and "I Understand Just How You Feel."

(Editor's Note: Forest entomologist Malcolm Furniss of Moscow, Idaho, researched the life of noted scientific illustrator Mary Foley Benson, 1905-1992. In her later years, she served as an illustrator for the UC Davis Department of Entomology.)
By Malcolm M. Furniss
Moscow, Idaho
MalFurniss@turbonet.com
My introduction to the work of scientific illustrator Mary Foley Benson (1905–1992) was happenstance. I am a forest entomologist, engaged for many years in research on bark beetles and associated organisms, including parasitoid Hymenoptera. Since 1963, I have resided in Moscow, Idaho, and have had ties with the former Bureau of Entomology laboratory at Coeur d'Alene, 90 miles to the north. There, Donald De Leon had studied Coeloides dendroctoni Cushman, a braconid preying on larvae of a scolytine, Dendroctonus ponderosae Hopkins, in western white pine. While reading his publication on the braconid's biology, I admired the drawing of a female wasp credited to Mary Foley (Fig. 1). A colleague noted my curiosity about this artist and called my attention to a photo of her in 1926 at age 21 (Fig. 2), when she was employed as a scientific illustrator with USDA. Subsequent inquiry led to discovering Mary's connections to entomology, including close acquaintance of the renowned Smithsonian Diptera taxonomist, John Merton Aldrich (1866–1934) (Fig. 3). I share here an account of Mary's life, concluding with her finding fulfillment and support in the community of Davis, Calif.
Mary Carilla Foley was born on April 2, 1905 at Storm Lake, Buena Vista County, Iowa, to Guy William Foley and Cora Helen Walrod. She had a brother, Charles, who had a flying service at Roosevelt Field, Long Island, and who had a pivotal role in her early adult life. Mary's inherent artistic talent became apparent while in first grade when her drawing of a puppy drew praise from her teacher (Wellings 1992). From then on, Mary made art her life and never forgot that teacher's encouragement.
At age 17, she left her Midwest roots to live with close family friends, John Aldrich and his wife, Della, in Washington, DC. Her intent was to go to art school—until then, she had been largely self-taught—but first, she had to earn tuition money. “Uncle Merton,” as she fondly referred to Aldrich, got her a job as an entomological draftsman with the USDA Bureau of Entomology. She learned quickly and eventually became Chief Scientific Illustrator. In evenings, she took art classes, graduating from the National School of Fine and Applied Arts and later attended the Corcoran School of Art. The closeness of Mary and Aldrich was like father and daughter. The tragic loss of his young son (Furniss 2010) may have affected his feelings for Mary, who would not have been much older.
Road Trip West with John and Della Aldrich, 1927
Mary and the Aldriches departed Washington on May 29, 1927 (nine days after Lindberg's Atlantic flight) in their Chevrolet (Fig. 4) coach loaded down with camping gear and headed across the continent on an adventure of their own. Aldrich kept a meticulous, detailed diary. A typed copy was discovered at the University of Idaho, Department of Entomology after his death and published by Paul Arnaud Jr. (Aldrich 2001). References to Mary appear throughout the diary and provide rare personal recollections of her. Her relatives and her closest friend, Margaret Hoyt, had passed away prior to my interest in her biography.
Driving was shared; however, it became apparent that Mary liked to drive and John often complimented her skill, so she drove the most. Two days into the trip, Aldrich noted: “Mary made fast time, keeping around 45.” Mary had little opportunity to paint until they reached the Rocky Mountains and a place named Wagon Tongue where Aldrich began collecting flies. Here she made water color paintings of a flower (Doedecatheon) and others not identified. Later, near Wells, Nevada, John packed up two Schmitt boxes pinned so far on the trip and mailed them with several small boxes of unmounted flies to Washington: “… some thousands of Diptera to be worked over later at home. Some good things there, I know.”
Eventually, they reached California and entered Yosemite National Park via Tioga Pass on July 4, having to drive over a patch of remnant snow. Nearing Bridal Veil, they came to a bear beside the road being fed by others. Mary stepped forward to get in a photo, but the bear was in the shade. Aldrich asked her to move the bear into the sun. As she came close to him, he gave her a bite above the knee; however, it only roughened the skin. Afterward, they made light of the episode.
Mary departed at Portland by train to return to her job. As John and Della drove up the scenic Columbia River highway heading for Moscow, they reminisced about her and missed having her company; they wished that she could be there at every turn. After thousands of miles cramped together, she had, indeed, become family!
John Aldrich had met and married Della Smith in Moscow after the untimely death of his first wife, Ellen Roe, and infant son, Spencer. The Aldriches were in Moscow from July 19 to Aug. 4, 1927, visiting Della's relatives and some of his acquaintances from his time at the University of Idaho, 1893–1913. On Sept. 2, the travelers reached home after being gone since 29 May. Aldrich died seven years later while completing plans to start early in June on another of his biennial collecting trips to the Pacific Coast. He was buried beside Ellen and son Spencer in the cemetery at Moscow.
Aviatrix Years
Mary's job at the Bureau of Entomology included painting insect-damaged crop plants such as appeared in the 1952 Yearbook of Agriculture (USDA 1952). Perhaps to break the boredom, she painted a parade of striped cucumber beetles across the office wall. Her boss was not amused (Wellings 1992). That phase of her life changed with marriage to Russell Benson, a patent lawyer and engineer with the US Patent Office, in 1928 and birth of a son, John, in 1931. In a Washington Post interview (Lewis 1937), she seemed to be living a normal life of a woman of the time “… keeping a home for her husband and small son, doing most of the cooking and making most of her own clothes.” They had designed their house in Georgian Colonial style with a studio for her painting. During the previous summer, she had laid a flagstone terrace while Russell built an outdoor fireplace. However, their relationship was less tranquil than it was portrayed.
Four years after her marriage, Mary had been given her first airplane ride by her brother, Charles, who had charge of one of the hangars at Roosevelt Field, Long Island (from where Lindberg had taken off on his Atlantic flight in 1927). That experience led to her taking flying lessons at the College Park, MD airport two or three times per week until receiving her pilot's license. Interviewed in later years, she divulged that she had kept her flying lessons secret from Russell (Wellings 1992). The marriage had become stressful and she found flying to be a great outlet from her exacting art work and her difficulties at home: “When I was flying, I left all my trouble on the ground. There was nobody up there but God and me.”
When Mary soloed in 1937, Charles gave her a J-2 Cub (Fig. 5). Her spirited nature showed in her flying: “I used to have such fun with it. Before I would land, I would wind it up with three giant loops in a row before they told me not to [not designed for aerobatics] (Haag 1983).” Owning an airplane expanded her involvement with flying. She became active in aviation organizations, including membership in The Ninety-Nines, an international organization for licensed women pilots founded by Amelia Earhart. She also joined the Washington Air Derby Association and placed second in her first air race, at College Park in May 1939 (Lewis 1937).
As World War II loomed, Mary's marriage was ending. Her son was in boarding school, so she joined Charles at his aviation business at Roosevelt Field, studied radio and navigation, became a member of the Civil Air Patrol and began ferrying airplanes for the War Training Service (Wellings 1992). She did not speak of her ferrying experiences except for saying that townsfolk came to see the woman flier after she was forced to land in a farm pasture and was waiting for a new propeller. She enlisted in the Women's Army Corps on Sept. 9, 1943 and instructed navigation to bomber crews in an on-ground Link Celestial Navigation Trainer. She regretted that women pilots were not allowed then to teach in the air.
When war ended, Mary relocated to Los Angeles where she attended Otis College of Art and Design from 1948–1951. Her training included instruction from Norman Rockwell, who spent the winter months there as artist-in-residence: “He was a wonderful person, with a fine sense of humor, and a scientific illustrator as well as a historian” (Tracy 1977). Afterward, she was a free-lance illustrator at Benson Studios. However, she did not attract the attention of newspaper reporters the way she had at Washington.
Home at Last … Davis
A great change came in 1964, when she was hired by Howard McKenzie at the University of California, Davis, as an illustrator for his monumental book on mealybugs (McKenzie 1967). She must have impressed him with her renditions of clearwing moths in Bulletin 190 of the U.S. National Museum (Engelhardt 1946) (Fig. 6). That publication contains 167 of her watercolor paintings of adult male and female clearwing moths and 87 line drawings of their genitalia. The paintings are eye-catching. When my nephew, Sean Furniss, saw them among the Engelhardt holdings at the Smithsonian, he was impressed with how real they appeared. I marvel that anyone could capture such natural colors by mixing pigment and rendering such life-like images with a brush.
Mary painted and drew mealybug illustrations for Howard McKenzie until his death in 1968. His 526-page treatise of California species has interspersed among its pages 21 of her watercolor paintings of species in their plant habitats (Fig. 7). The book also includes many of her line drawings of microscope slides of holotype adult females, split longitudinally by dorsal and ventral views (Fig. 8). The drawings include minute detail only seen by microscope at high magnification. Mary had become skilled in microscopy and, although not formally trained in entomology, she had to have learned much technical detail about what she was asked to draw. After McKenzie died, Douglass Miller completed two joint manuscripts containing illustrations by Mary (Miller and McKenzie 1971, 1973). Earlier, she had illustrated mealybugs in the monograph on the genus Asterolecanium by Russell (1941).
Announcement of publication of McKenzie's book noted that it had been illustrated by Mary and included her pedigree dating to her employment with USDA Bureau of Entomology in Washington DC during 1922-1928 as Chief Scientific Illustrator. Her presence in the community had already become known by her participation in Chamber of Commerce events, teaching painting techniques through the Davis adult education program and scientific illustration through the University of California Cooperative Extension.
In the years after McKenzie's death, she contracted a painting project for Harry Lange to illustrate a book on insects infesting California crops. The idea was to show both the insect and its host plant in realistic fashion. However, when Mary portrayed the plant, the accompanying insect was at the same scale, much too little to suit entomologist Lange. His wife, Ellen Lange, recalls that, “A real problem with depicting insects and plants is the difference in size. Harry Lange used to have battles with Mary about the plants being too big, i.e., emphasis on the plants versus the insects.” They resolved the problem by superimposing a “magnifying glass” to enlarge the insect (Fig. 9). The proposed publication never materialized; her 55 paintings are stored in the Bohart Museum of Entomology at Davis. In looking back, such paintings (and to some extent, drawings) were being replaced by digital photography and scanning electron microscopy.
Having retired from the Department of Entomology work, Mary (Fig. 10) said: “For the first time in my life, I can paint what I feel like painting most. I have always wanted to paint wildflowers, especially California wildflowers.” (Barr, 1992). She set about doing just that and put her paintings on display for sale at public places, gradually developing a devoted following. “This is the greatest place in the world for me. I get such support from the people of Davis. They hang my work all over town.” (Haag 1983).
An event in 1983 crowned her rise to prominence beyond the Davis community. At the Smithsonian's invitation, 45 of her paintings were displayed there during May-June, entitled, “Paintings of California Flora.” In her letter of Nov. 20, 1982 to Officer of Exhibits, William Haase, she listed the paintings, some of which included insects, that were to be sent and suggested the name California Flora for the exhibit. She singled out two of the paintings that merited special attention: “The Golden Lupine (Fig. 11) which grows around Davis, along the railroad track. It is almost extinct in the wild. There is some talk of making it the Davis city flower.” (She subsequently donated the original watercolor painting to the city.) And, “The second painting is the California poppy and wild oats. I had hoped to give it to our First Lady, Nancy Reagan, but could not decide just how to go about it. It seemed appropriate since the poppy is our state flower.” The letter concluded with a question regarding who should print the brochure for the show and that the paintings would be shipped by truck. She requested that, after the show, they should be sent to her residence at 1408 Claremont Drive, Davis. No one whom I contacted knew of the fate of these paintings. However, she evidently had prints made of some for sale. I noted one of her paintings being resold by Valley Auction on the Internet; it was number 123 of 500, entitled, “Indian Paint Brush and Fiddleback,” gathered from a vacant lot near her home. It portrays her eloquent, simplistic style that is so appealing to me.
Her dream of a Smithsonian exhibit having come true was timely. Her eyesight began failing soon after. She kept on painting but results were more abstract. During those challenging final years, she kept a positive outlook. As recalled in her obituary (Barr 1982) she said, “Life is like a stream; if you reach a stone you go around it. So now, I'm moving on after the stone.” Mary died at Davis on 18 June 1992 at age 87.
Acknowledgments
Sandra Kegley, Coeur d'Alene, ID, provided Fig. 2, which identified Mary and thereby initiated my interest in pursuing this article. Sean and Martha Furniss, Reston, VA, provided newspaper articles and searched the Smithsonian archives for information about Mary. Ellen Lange, Davis, CA, provided personal recollections of Mary and of her paintings for Harry Lange. Lynn Kimsey, Bohart Museum of Entomology, University of California, Davis, provided obituaries of Mary and a listing of Mary's paintings stored there. Amy Thompson, Special Collections and Archives Library, University of Idaho, provided the photo of Aldrich. Luc Leblanc, W.F. Barr Entomological Museum, University of Idaho, loaned references concerning Aldrich and Mary from the museum library. The manuscript was reviewed by Sandra Kegley, Ellen Lange, Linwood Laughy, and Douglass R. Miller, Systematic Entomology Laboratory, USDA, Beltsville, MD.
Literature Cited
Aldrich, J. M. 2001. Diary of a Western Trip, 1927. Myia - A publication on entomology 6:235-301. California Academy of Sciences.
Barr, P. 1992. Prominent Davis Artist Dies. Davis Enterprise. June 22.
De Leon, D. 1934. The Morphology of Coeloides dendroctoni Cushman (Hymenoptera: Braconidae). Journal New York Entomological Society. XLII: 297-317.
Engelhardt, G. P. 1946. The North American Clear-Wing Moths of the Family Aegeriidae. Bulletin 90. U. S. National Museum. 222 pp.
Furniss, M. M. 2010. John Merton Aldrich (1866-1934) – A Forest Entomologist's View of Idaho's Renowned Fly Taxonomist. Latah Legacy 38:18-23.
Haag, J. 1983. A Californian's Floral Fantasy. The Sacramento Bee. July 24.
Lewis, R. 1937. Vocation and Avocation Combined by D. C. Artist. The Washington Post. October 17.
McKenzie, H. L. 1967. Mealybugs of California with Taxonomy, Biology and Control of North American Species (Homoptera: Coccoidea: Pseudococcidae). University of California Press, Berkeley. 525 pp.
Miller D. R., and H. L. McKenzie. 1971. Sixth Taxonomic Study of North American Mealybugs with Additional Species from South America. Hilgardia. 40:565-602.
Miller, D. R., and H. L. McKenzie, 1973. Seventh Taxonomic Study of North American Mealybugs (Homoptera: Coccoidea: Pseudococcidae). Hilgardia 41: 489-542.
Russell, L. M. 1941. A Classification of the Scale Insect Genus Asterolecanium. U.S. Department of Agriculture, Miscellaneous Publications 424: 1-319
Tracy R. L. 1977. Mary Foley Benson. Her Wildflower Paintings Are “Expression of Gratitude” The Sacramento Bee. August 20, 1977.
USDA 1952. The Yearbook of Agriculture. U.S. Government Printing Office.
Wellings, M. 1992. Davis Has Lost a Free Spirit. The Davis Enterprise. June 23.
- Author: Kathy Keatley Garvey
Dr. Mussen, a resident of Davis, was admitted to a local hospital on May 25. He was diagnosed with liver cancer/failure on May 31 and returned to the family home June 1 for hospice care. He passed away the evening of June 3.
“Eric was a giant in the field of apiculture,” said Steve Nadler, professor and chair of the UC Davis Department of Entomology and Nematology. “The impact of his work stretched far beyond California.”
Dr. Mussen, known to all as “Eric,” joined the UC Davis entomology department in 1976. Although he retired in 2014, he continued his many activities until a few weeks prior to his death. For nearly four decades, he drew praise as “the honey bee guru,” “the pulse of the bee industry" and as "the go-to person" when consumers, scientists, researchers, students, and the news media sought answers about honey bees.
“Eric's passing is a huge loss," said longtime colleague Lynn Kimsey, director of the Bohart Museum of Entomology and a UC Davis distinguished professor of entomology. "He was always the go-to person for all things honey bee. He worked happily with hobbyists, commercial beekeepers and anyone just generally interested."
Colleagues described Mussen as the “premier authority on bees and pollination in California, and one of the top beekeeping authorities nationwide,” “a treasure to the beekeeping industry," and "a walking encyclopedia when it comes to honey bees.”
“Eric's career was so productive and exciting that a book would be required to do justice for his many contributions to his profession as extension entomologist specializing in apiculture, better known as beekeeping,” Gary said. “His mission basically was facilitating productive and reciprocal communication between beekeeping researchers at UC Davis, commercial beekeeping as it affects California's vast needs for the pollination of agricultural crops, providing helpful information to hobby beekeepers, and educating the general public concerning honey bees. His great professional successes in all areas have been recognized around the world. He has received numerous awards, especially from the beekeeping industry. He was by far the best Extension apiculturist in this country!”
“In addition to professional duties, he enthusiastically tackled other projects for entomology faculty,” Gary said. “For example, he critically reviewed most of my publications, including scientific papers, books, and bulletins. He worked diligently to help create the Western Apicultural Society and later served as president. (Mussen served six terms as president, the last term in 2017.) I especially appreciated his volunteering to moderate a video that historically summarized and recorded my entire 32-year career at UC Davis. And his tribute would not be complete without mentioning that he was one of my favorite fishing buddies.”
“Eric was proud of his loving wife, Helen, and family (sons Timothy and Christopher and two grandchildren),” Gary said. “His family support was unconditional. He will be sadly missed by everyone.”
Leslie Saul-Gershenz, a UC Davis doctoral alumnus in entomology, praised his kindness and generosity. “I have known Eric for over 30 years and he was the kindest, most supportive human being. He always came to speak at the San Francisco Bee Club to support local beekeepers and was the most generous of colleagues anyone could ever hope for. He was a immense resource of knowledge about honey bees, and I am sure he will be missed by many people across the state of California.”
Mussen's longtime friends and colleagues--bee breeder-geneticist Susan Cobey of Washington State University (WSU), a former manager of the Harry H. Laidlaw Jr. Honey Bee Facility, UC Davis, and her husband, Timothy Lawrence, WSU associate professor and county director, WSU Extension, Island County--are heartbroken. “Eric is an icon of the beekeeping industry and beyond, a career shaper, problem solver, the information man who always had an answer or would find one, and, always given with integrity, regardless of the issue, biological or political, to whoever posed the question and need," Cobey said. "His contributions, impact and love from the people he touched will live, continuing to contribute and benefit their lives. His spirit is with us.”
Jackie Park-Burris of Jackie Park-Burris Queens, Inc., Palo Cedro, a leader in the queen bee breeding and beekeeping industries, said she met Eric more than 40 years ago “and from day one he was mentoring me. He was the bee guy for the entire country! Eric was the bee industry's connection to the scientific world. Eric understood both camps and he connected them. Eric had incredible integrity that I have never seen matched. Because of that integrity, beekeepers felt confident in sharing their problems with him, knowing their secrets were safe. Eric always voiced the opinion he felt was right, even if it wasn't the most popular.”
“Eric told me that he looked at the bee industry as his family,” said Park-Burris, a past president of CBSA and a member of the board of directors for more than 20 years. “When my son attended UC Davis, he and Helen made sure Ryan knew he could contact them if he needed anything. Eric even came to a function on campus that my son was in charge of to show support.”
“At last year's CSBA convention. I was awarded the Eric Mussen Distinguished Service Award. Eric sent me an email, congratulating me and told me he could not think of a more qualified person to receive it. It bought a tear to my eye back then, now I will treasure that email even more.” A photo of an early-career Mussen appears on her website.
Patrick “Pat” Heitkam, owner of Heitkam's Honey Bees, Orland, recalled the beekeeping industry's turbulent times in the 1980s. “He was many things and I'm certain no one will fill his position as admirably as he did. The mid-'80s were a turbulent time for commercial beekeepers in California. The advent of tracheal mites and later varroa disrupted our ability to make a living. We were affected in different ways. Passions ran high. Eric maintained the respect of all concerned. I was frustrated, more than once, because he wouldn't tell me what I wanted to hear. He was a scientist, stayed neutral, and only commented when he could be factual. I saw him as the level-headed patriarch of his dysfunctional beekeeping family. This is one of his many attributes. I can't say enough about his value to bee world.”
Randy Oliver of ScientificBeekeeping.com ("beekeeping through the eyes of a biologist") treasured him. "Eric Mussen was not only a longtime friend and collaborator of mine, but a model for me. He was a beloved and exemplary Extension apiculturist, with his engaging presentations, interpreting the science for the benefit of beekeepers. His monthly newsletter was an important source of information to not only California beekeepers, but also to many across the United States and around the world. Eric always made time to happily share information with anyone who asked. I reminisced with his wife Helen shortly before his passing about his joy at doo-wop singing, and I'm sure that that's how this bright light of a man would like to be remembered."
Amina Harris, director of the UC Davis Honey and Pollination Center and co-owner of Z Specialty Food, Inc., commented that "When I first came to California in the early 1980s, I was working with my husband establishing and growing a varietal honey company. One of the first people I met at UC Davis was Eric Mussen, the state apiculturist. Eric was someone who had a lot of answers to a lot of questions! Ever the educator, Eric was well versed in all of the issues of the bee world and readily shared his knowledge with any and everyone who asked. His answers were always down to earth and understandable, with his wry Midwestern sense of humor running underneath. You'd ask a question – and you always got an answer!"
Born May 12, 1944 in Schenectady, N.Y., Mussen received his bachelor's degree in entomology from the University of Massachusetts (after declining an offer to play football at Harvard) and then obtained his master's degree and doctorate in entomology from the University of Minnesota in 1969 and 1975, respectively. Mussen credits his grandfather with sparking his interest in insects. His grandfather, a self-taught naturalist, would take his young grandson to the woods to point out flora and fauna.
Bees became his life, and Mussen thoroughly enjoyed his career. For nearly four decades, Mussen wrote and published the bimonthly newsletter, from the UC Apiaries, and short, topical articles called Bee Briefs, providing beekeepers with practical information on all aspects of beekeeping. His research focused on managing honey bees and wild bees for maximum field production, while minimizing pesticide damage to pollinator populations.
A favorite among the news media for his bee expertise and personality, Mussen appeared on the Lehrer Hour, BBC, Good Morning America, National Public Radio (Science Friday) and other major news outlets, and was quoted by the New York Times, Boston Globe, and Los Angeles Times, among others.
Highly honored by his peers, Mussen received the 2006 California Beekeeper of the Year award; the American Association of Professional Apiculturists' 2007 Award of Excellence in Extension Apiculture; the 2008 Distinguished Achievement Award in Extension from the Pacific Branch of the Entomological Society of America; the 2010 statewide Pedro Ilic Outstanding Agricultural Educator; and was a member of the UC Davis Bee Team that won the 2013 team award from the Pacific Branch of the Entomological Society of America.
Among his scores of awards: the 2013-14 Distinguished Service Award for Outstanding Extension from the UC Agriculture and Natural Resources (UC ANR) and the 2013 Alexander Hodson Graduate Alumni Award from his alma mater, the University of Minnesota.
Gene Brandi, the 2018 president of the American Beekeeping Federation, remembers presenting him with the prestigious Founders' Award from the Foundation for the Preservation of Honey Bees at the 75th annual American Beekeeping Federation conference in Reno. "Eric received a well-deserved rousing standing ovation!” said Brandi, extolling him as "an outstanding liaison between the academic world of apiculture and real-world beekeeping and crop pollination."
"Eric was a legend in the beekeeping world who was always willing to go the extra mile to help beekeepers and bee industry organizations deal with issues pertaining to honey bee health, regulations, and various threats to the industry," Brandi said. "He also helped agricultural organizations, government officials, and the general public better understand the value of honey bees to the world. Eric was a great advocate for the honey bee and beekeepers. He was truly a national treasure.
"Eric served on the California State Beekeepers Association Board of Directors for 39 years as apiculturist, but he also was the parliamentarian for many years, and due to his long tenure, was quite the historian as well," Brandi added. "He co-founded the Western Apicultural Society and served as president for six terms. He founded the American Association of Professional Apiculturists and was either president of secretary/treasurer for the first ten years."
"In addition to his work with the aforementioned organizations, Eric was the UC Honey Bee Liaison to California Department of Food and Agriculture, California Department of Pesticide Regulation, EPA, California Farm Bureau Federation, Almond Board of California, National Honey Board, California Bee Breeders' Association, and others. Eric was always willing to help the beekeeping industry by testifying at hearings or writing letters to support the bee industry on various issues. He received many awards from a number of organizations, but the ones with which I am most familiar are the California State Beekeepers Association Distinguished Service Award as well as the Beekeeper of the Year Award. The Foundation for the Preservation of Honey Bees honored Eric in 2018 with the coveted Founders; Award.
Brandi related that "Eric also conducted a number of research projects over the years including research on sacbrood virus, effects of medfly sprays on honey bees, effects of antibiotics of honey bee brood, potential of Neem as a varroa control, effects of selected fungicides on honey bee brood, effects of antibiotics, and effects of high-fructose corn syrup on packaged bee development. Eric's "From the UC Apiaries" newsletter was a renowned publication which contained valuable information about current events in the beekeeping world which beekeepers needed to know. There is so much more that can be said about Eric and his many contributions to the beekeeping world."
When Mussen was nominated (and received) the 2013-14 Distinguished Service Award for Outstanding Extension from the UC Agriculture and Natural Resources (UC ANR), MacArthur Genus Awardee Professor Marla Spivak, Distinguished McKnight University Professor Apiculture/Social Insects at the University of Minnesota, wrote: "Without question, Eric is the No. 1 Extension person dealing with honey bees in the nation, if not the world. Research colleagues, beekeepers and the public are all very lucky to have him.”
Mussen's team of nominators singled him out for his “amazing knowledge of bees, his excellent communication skills in a diverse clientele, including researchers, Extension personnel, legislators, commodity boards, grower organizations, pesticide regulators, students, news media, and beekeeping associations at the national, state and local levels; his eagerness to help everyone, no matter the age or stature or expertise, from an inquiring 4-H'er to a beginning beekeeper to a commercial beekeeper; and his ability to translate complicated research in lay terms.” He presented multiple talks to every beekeeping association in California, and spoke at a variety of events, including pollinator workshops, animal biology classes, UC events, and fairs and festivals.
The nominators lauded him for “his valuable research, which includes papers on antibiotics to control American foulbrood; fungicide toxicity in the almond orchards; the effect of light brown apple moth mating pheromone on honey bees; the effects of high fructose corn syrup and probiotics on bee colonies; and the invasion and behavior of Africanized bees. He was often consulted on colony collapse disorder and bee nutrition.”
Overall, Mussen was known as "a well-respected, well-liked, honest, and unflappable person with a delightful sense of humor."
Valeri Strachan-Severson of Strachan Apiaries, Inc., Yuba City, producer of New World Carniolan queen bees, treasured his advice. "After my dad, Don Strachan, passed away, I made a pest of myself contacting Eric for advice. He never made me feel like I was asking dumb questions and always had the answers. Sometimes he'd say he'll get back to me if he wasn't sure, and he always did. He was an encyclopedia of apiary knowledge. I'll miss him. He was seriously one of a kind."
Mussen was a longtime board member of the California State Beekeepers' Association and a consultant for the Almond Board of California. He co-founded the Western Apicultural Society, serving six terms as president, the last during the 40th anniversary conference at UC Davis in 2017. He also was involved in the formation of the American Association of Professional Apiculturists and held the offices of president or treasurer of that association for many years. He was a scientific advisor to the UC Davis Honey and Pollination Center.
Mussen was instrumental in the development of the Almond Board of California's Honey Bee Best Management Practices for Almonds. The Almond Board honored him with a service award, describing him as being an “authoritative and trusted source for guidance on research, technical, and practical problem solving and issues facing both industries.” Shortly after he retired, both the CSBA and WAS created an Eric Mussen Award to present to their outstanding members.
“Eric is a worldwide authority on honey bees, but no problem is too small and no question too involved for him to answer,” said the late Extension entomologist Larry Godfrey prior to Mussen's retirement. “He devotes his research and extension activities to the improvement of honey bee health and honey bee colony management practices. Eric helps growers, consumers, UC Farm Advisors, agricultural commissioners, scientists, beekeepers, researchers, pesticide regulators, 4-H'ers, and state and national agricultural and apicultural organizations. He ignites their interest in maintaining the health of bees, cultivates their friendship, and generously gives of his time and intellect.”
William Hutchison, professor and head of the Department of Entomology at the University of Minnesota, who presented him with the 2013 Alexander Hodson Graduate Alumni Award, related at the ceremony that Mussen tackled "many new challenges–mites, diseases, and Africanized honey bees, to name a few–to enhance the pollination success of California's diverse agricultural cropping systems, with considerable emphasis on almonds. In brief, he is in demand, and he continues to be a primary source for objective information on honey bee health and pollination in California."
In his retirement years--in between his family commitments and his hobbies of birding and singing doo-wop with a local group--Mussen not only served as the 2017 president of the Western Apicultural Society but doubled as the program chair at the UC Davis conference. He reviewed funding proposals for Project Apis m., which makes funding decisions and handles the funds for the National Honey Board and other entities; and served on the scientific review panel for the Bee Informed Partnership organization, which reviews funding requests of tech teams. He was a scientific advisor to the UC Davis Honey and Pollination Center.
"I am basically all pro-bee,” Mussen told the American Bee Journal in a two-part feature story published in September 2011. “Whatever I can do for bees, I do it...It doesn't matter whether there is one hive in the backyard or 15,000 colonies. Bees are bees and the bees' needs are the bees' needs.”
Celebration of Life. A Celebration of Life is planned from 4 to 6 p.m., Sunday, Aug. 28 in the Putah Creek Lodge, UC Davis campus. RSVP here. For those unable to attend, a live webinar will be produced by UC Davis distinguished professor Walter Leal of the Department of Molecular and Cellular Biology, and a former chair of the Department of Entomology. Registration is underway here at https://bit.ly/3czl5Am. It also will be on YouTube.
Family and friends suggest memorial contributions be made to the California State 4-H Beekeeping Program, with a note, "Eric Mussen Memorial Fund." Mary Ciriceillo, director of development for the UC Agriculture and Natural Resources, said checks may be made out to the California 4-H Foundation and mailed to:








