- Author: Kathy Keatley Garvey
Boudinot, who studied with major professor Phil Ward of the UC Davis Department of Entomology and Nematology, is the second UC Davis-affiliated entomologist to receive the honor in its 28-year history. Jessica Gillung of the Lynn Kimsey lab, Bohart Museum of Entomology, won the award in 2019.
Snodgrass (1875-1962), a leader in insect morphology, is known for his 76 scientific articles and six books, including Insects, Their Ways and Means of Living (1930) and the book considered to be his crowning achievement, the Principles of Insect Morphology (1935).
Boudinot has them all. “Principles of Insect Morphology and the Anatomy of the Honey Bee were the foundation of my studies,” he said. “I have both, plus his Textbook of Arthropod Morphology and Insects, Their Ways and Means of Living on my desk in the lab.”
The Snodgrass Award, which includes a certificate and cash prize, recognizes outstanding research by a PhD student who has completed a research thesis or dissertation in arthropod morphology, systematics, taxonomy, or evolution. Nominees are scored on honors, awards, achievements and recognition; recommendations of professors and advisors; grantsmanship, publications, creativity and innovation of thesis or dissertation; and contribution to morphology.
Boudinot's dissertation: “Systematic and Evolutionary Morphology: Case Studies on Formicidae, Mesozoic Aculeata, and Hexapodan Genitalia.”
He earlier received the prestigious Alexander von Humboldt Research Fellowship to do research on evolutionary and comparative anatomy in Jena, Germany. He will locate to Germany in early 2021 for the two-year fellowship, after completing intensive language studies.
'I Am a Morphologist Because of Robert Evans Snodgrass'
“I am a morphologist because of Robert Evans Snodgrass,” Boudinot wrote in his application. “Although I have had pressure from my earliest undergraduate to become a molecular systematist, it was my chance encounter with Snodgrass's Principles of Insect Morphology late one night in the college library that set the course of my career. I had struggled for years at that point to understand the biodiversity of insects and to untangle the deep morass of arcane terminology, but my vision was illuminated by the conceptual clarity, linguistic simplicity, and exceptional engravings of the Principles. This work continues to hold special dominion over my thinking, as it is through the principle of mechanical function for explaining comparative anatomical observations that I have come to my present understanding of systematic and evolutionary morphology.”
Boudinot wrote that his “career objective, in brief, is to resolve the morphological evolution of insects through the synthesis traditional morphology, as represented by Snodgrass, with recent trends in digital anatomy and bioinformatics. I envision a future wherein students rely not on Borror & Delong, a holdover from Comstock's 19thCentury manual, but rather learn about insect structure, function, classification, and evolution through manipulation of three- and four-dimensional digital anatomical models, substantiated via manual curation and dissection. I want students to see for themselves what I have understood through the study of Snodgrass's work, balanced by contemporary research: The origin of the Arthropoda and morphological transformation through geological time to the resplendent, and endangered, diversity of today.
“In sum, my identity as an entomologist, and as a scientist more broadly, is due to the insights on the language and phenomenology of morphological evolution I gained from the oeuvre of Snodgrass. Without these works, I would probably still be a botanist.”
Boudinot's research interests include the origin and evolution of complex phenotypic systems from the perspective of phylogenetic systematics, including molecular and paleontological evidence. Specializing on morphological identity and evolutionary transformation, the skeletomuscular system of Arthropoda, with emphasis on the male genitalia of Hexapoda and systematics of the Hymenoptera, particularly the Formicidae.
John Henry Comstock Award
Highly honored for his academics, leadership, public service activities, professional activities and publications while at UC Davis, Boudinot received the 2019 John Henry Comstock Award, the top graduate student award given by ESA's Pacific Branch. The branch encompasses 11 Western states, U.S. territories, and parts of Canada and Mexico.
In the Comstock award application, Steve Nadler, professor and chair of the Department of Entomology and Nematology, described Boudinot as “A highly respected scientist, teacher and leader with a keen intellect, unbridled enthusiasm, and an incredible penchant for public service.” Boudinot maintained a 4.00 grade point average and published 18 peer-reviewed publications on insect systematics, some landmark or groundbreaking work.
His most recent publications: one on Cretaceous Strepsiptera in Cladistics and the other on the iron maiden ants in Myrmecological News ("Two New Iron Maiden Ants from Burmese Amber (Hymenoptera:
Formicidae: †Zigrasimeciini").
Boudinot received multiple “President's Prize” awards for his research presentations at national ESA meetings. He organized the ESA symposium, “Evolutionary and Phylogenetic Morphology,” at the 2018 meeting in Vancouver, B.C. , and delivered a presentation on “Male Ants: Past, Present and Prospects” at the 2016 International Congress of Entomology meeting in Orlando, Fla.
Boudinot served on—and anchored—three of the UC Davis Linnaean Games teams that won national or international ESA championships. The Linnaean Games, now known as the Entomology Games, are a lively question-and-answer, college bowl-style competitions on entomological facts played between university-sponsored student teams.
Brendon served as president of the UC Davis Entomology Graduate Student Association from 2006 to 2019, and co-chaired the department's UC Davis Picnic Day celebration (with forensic entomologist Robert Kimsey) for three years.
Before enrolling in graduate school program at UC Davis in 2014, Brendon received his bachelor's degree in entomology in 2012 from The Evergreen State College, Olympia, Wash. Professor John T. Longino served as his mentor.
- Author: Kathy Keatley Garvey
Boudinot presented a talk in Germany at the 9th Dresden Meeting on Insect Phylogeny, hosted by the Senckenberg Institute. In his presentation, "Skeletomuscular Evolution of Male Insect Genitalia, with Emphasis on the Endopterygota," he explained the second chapter of his thesis to an international audience of entomologists, spanning phylogeneticists to anatomists.
At the XXIV Simpósio de Mirmecologia in Belo Horizonte, Minas Gerais, Brazil, Boudinot will present preliminary results from his third and final thesis chapter in a symposium titled "Morphology Reloaded: Conquering the Third Dimension of Ant Evolution and Biogeography." His talk is titled "Altered States: Aculeate Anatomy and the Early Evolution of the Formicoidea."
"While this symposium is focused on the application of scanning technologies, such as X-ray computed microtomography, I will be speaking about the evolution of ant morphology through evolutionary time," Boudinot related. He will be presenting results from combined phylogenetic analysis of genome-scale DNA sequence data and a massive dataset of 576 morphological characters scored for 573 species.
"One of the novel aspects of my study is how many fossils I have included (more than 300)," Boudinot said. "The objective of this particular work is to estimate the geological age in which the ants originated, as well as the physical features of the ancestral ant, which I will then use to trace evolutionary and paleoecological patterns to the modern fauna."
Boudinot received the prestigious John Henry Comstock Award in April at the Pacific Branch, Entomological Society of America (PBESA) meeting in San Diego. This is the highest graduate student award given by PBESA, which encompasses 11 Western states, U.S. territories, and parts of Canada and Mexico. Its parent organization, the Entomological Society of America (ESA), will honor the six Comstock award winners--one from each branch--at its Nov. 17-20 meeting in St. Louis.
Boudinot, who was praised for his academic record, leadership, public service activities, participation in professional activities, and his publications, is the sixth UC Davis recipient of the Comstock award:
2019: Brendon Boudinot (major professor Phil Ward)
2015: Mohammad-Amir Aghaee (major professor the late Larry Godfrey)
2014: Kelly Hamby (major professor Frank Zalom)
2013: Matan Shelomi (major professor Lynn Kimsey)
2008: Christopher Barker (major professor William Reisen)
1983: Elaine Backus (major professor the late Donald McLean)
Active in PBESA and ESA, Boudinot received multiple “President's Prize” awards for his research presentations at national ESA meetings. He organized the ESA symposium, “Evolutionary and Phylogenetic Morphology,” at the 2018 meeting in Vancouver, B.C. , and delivered a presentation on “Male Ants: Past, Present and Prospects” at the 2016 International Congress of Entomology meeting in Orlando, Fla.
Boudinot served on—and anchored—three of the UC Davis Linnaean Games teams that won national or international ESA championships. The Linnaean Games are a lively question-and-answer, college bowl-style competition on entomological facts played between university-sponsored student teams.
Boudinot has served as president of the UC Davis Entomology Graduate Student Association since 2006, and is active in the campuswide UC Davis Picnic Day; he has co-chaired the department's Picnic Day Committee since 2017.
- Author: Kathy Keatley Garvey
The award, given by ESA's Systematics, Evolution and Biodiversity Section, recognizes outstanding research by a doctoral student who has completed a research thesis or dissertation in arthropod morphology, systematics, taxonomy, or evolution. The prize is a $500 cash award and an invited talk at ESA. She'll speak on "Unraveling the Evolution of Spider flies (Diptera, Acroceridae): Progress and Possibilities" at the ESA's annual meeting, set Nov. 17-20 in St. Louis, Mo.
Gillung is the first UC Davis student to receive the award, which was first presented in 1992. Her dissertation, “Systematics and Phylogenomics of Spider Flies (Diptera, Acroceridae),” focused on the evolution, conservation, biology, and taxonomy of spider flies, a group of spider natural enemies.
Awarded her doctorate in entomology in December 2018, Gilllung studied with major professor Lynn Kimsey, director of the Bohart Museum of Entomology and professor of entomology, UC Davis Department of Entomology and Nematology. She also studied with mentor Shaun Winterton, insect biosystematist, California Department of Food and Agriculture, and collaborated with ant specialist and taxonomist Phil Ward, professor, UC Davis Department of Entomology and Nematology.
Gillung's dissertation, involving genomics, phylogenetics, systematics, and comparative analyses, “has increased our understanding of the biological patterns and processes that have shaped our planet's biodiversity,” Kimsey wrote in her letter of nomination. Her taxonomic work included identification keys and morphology-based diagnoses of species using modern techniques of cybertaxonomy—the application of the internet, digital technologies, and computer resources to increase and speed up the discovery and cataloging of new species, Kimsey wrote in her letter of nomination.
“Using cybertaxonomic tools, Jessica described 25 new spider fly species herself, and in collaboration with fellow entomologists, three fossil species from Baltic amber, described in her first dissertation chapter," Kimsey wrote. "Cybertaxonomy is a powerful tool that allows researchers and citizen scientists to collaborate in real time and across great distances to increase the speed and efficiency of biodiversity discovery.”
Kimsey noted that “Jessica unraveled the functional and ecological implications of key morphological traits, as well as their distribution across the Tree of Life," and "established new homologies for the wing venation of spider flies. She conducted detailed and assiduous dissections of male reproductive structures (i.e., genitalia) to understand homologies, demonstrating that morphological traits are dynamically evolving systems useful for both classification and inference of evolutionary history.”
While at UC Davis, Gillung drew more $120,000 in grants and awards for her multifaceted research on genomics, bioinformatics, phylogenetics, plant-pollinator interactions, and biodiversity discovery. She compiled a near straight-A academic record, published 11 refereed publications in top journals, and engaged in public service and outreach programs that reached more than 20,000 people at UC Davis-based events.
Gillung was a key member of the UC Davis Linnaean Games Team that won the ESA national championship in 2015. The Linnaean Games are lively question-and-answer, college bowl-style competitions on entomological facts played between university-sponsored student teams.
The UC Davis-trained entomologist earlier received
- The 2019 Marsh Award for Early Career Entomologist, sponsored by the Royal Entomological Society. That involved a $1624 cash award and an invitation to the society meeting, Aug. 20-22 at the London School of Hygiene and Tropical Medicine.
- The 2019 Early Career Award from the Pacific Branch, Entomological Society of America (PBESA). PBESA encompasses 11 western states, U.S. territories and parts of Canada and Mexico.
- The 2018 Student Leadership Award from PBESA
- The “Best Student Presentation Award” at the ninth annual International Congress of Dipterology, held in 2018 in Windhoek, Namibia.
At Cornell, Gillung is researching Apoidea (stinging wasps and bees) phylogenomics, evolution and diversification in the Brian Danforth lab.
- Author: Kathy Keatley Garvey
She is the first UC Davis-affiliated scientist to win the award. She will receive a certificate, 1250 pounds ($1,624) and an expense paid trip to London to receive the award at the Ento 19 conference, set Aug. 20-22 at the London School of Hygiene and Tropical Medicine.
The Royal Entomological Society, an international organization devoted to the study of insects, was founded in 1833 as the Entomological Society of London. Its mission is to disseminate information about insects and improve communication between entomologists.
Gillung's work on spider flies, involving genomics, phylogenetics, systematics, and comparative analyses, “has increased our understanding of the biological patterns and processes that have shaped our planet's biodiversity,” wrote her major professor and award nominator Lynn Kimsey, director of the Bohart Museum of Entomology and a professor of entomology, UC Davis Department of Entomology and Nematology.
Gillung received her doctorate in entomology from UC Davis in December 2018, studying with Kimsey and mentor Shaun Winterton, insect biosystematist, California Department of Food and Agriculture, and a member of the Royal Entomological Society. Gillung also collaborated with ant specialist and taxonomist Phil Ward, professor, UC Davis Department of Entomology and Nematology.
Gillung is now a postdoctoral associate at Cornell University in the Bryan Danforth lab, where she is researching Apoidea (stinging wasps and bees) phylogenomics, evolution and diversification.
Her dissertation, “Systematics and Phylogenomics of Spider Flies (Diptera, Acroceridae),” focused on the evolution, conservation, biology, and taxonomy of spider flies, a group of spider natural enemies,
Gillung's taxonomic work on spider flies, described as landmark, included identification keys and morphology-based diagnoses of species using modern techniques of cybertaxonomy—the application of the internet, digital technologies, and computer resources to increase and speed up the discovery and cataloging of new species, Kimsey wrote. “Using cybertaxonomic tools, Jessica described 25 new spider fly species herself, and in collaboration with fellow entomologists, three fossil species from Baltic amber, described in her first dissertation chapter. Cybertaxonomy is a powerful tool that allows researchers and citizen scientists to collaborate in real time and across great distances to increase the speed and efficiency of biodiversity discovery.”
“Jessica unraveled the functional and ecological implications of key morphological traits, as well as their distribution across the Tree of Life,” Kimsey said. “In her doctoral dissertation, she established new homologies for the wing venation of spider flies and conducted detailed and assiduous dissections of male reproductive structures (i.e., genitalia) to understand homologies, demonstrating that morphological traits are dynamically evolving systems useful for both classification and inference of evolutionary history.”
Since many insect species are threatened, geographically restricted, or relatively rare in nature, Gillung performed non-destructive DNA extraction of specimens housed in entomological collections, including the Bohart Museum of Entomology. Gillung collected molecular data from decades-old insects without damaging the specimens.
Gillung's multifaceted research on genomics, bioinformatics, phylogenetics, plant-pollinator interactions, and biodiversity discovery drew more $120,000 in grants and awards while at UC Davis.
The UC Davis alumnus is known for her “phenomenal leadership activities, her nearly straight-A academic record (3.91 grade point average), her excellence as an entomologist and teacher, her public service and outreach programs (from 2013 to 2018, she reached more than 20,000 people at UC Davis-based events) and her incredible publication record,
Kimsey said. “She published 11 refereed publications related to her thesis in very strong journals. Most entomologists do not publish nearly that much, even as a postdoctoral scholar or a junior faculty member.”
A recipient of numerous other awards, Gillung won the prestigious international award for “Best Student Presentation Award” at the ninth annual International Congress of Dipterology, held in 2018 in Windhoek, Namibia. She also won the 2019 Early Career Award from the Pacific Branch, Entomological Society of America (PBESA) and the 2018 PBESA Student Leadership Award. PBESA encompasses 11 western states, U.S. territories and parts of Canada and Mexico.
Gillung was a key member of the 2015 PBESA championship Linnaean Team that went on to win the ESA national championship. The Linnaean Games are lively question-and-answer, college bowl-style competition on entomological facts played between university-sponsored student teams.
Gillung also collaborated on a project aimed at encouraging students to attend and participate in the Orlando, Fla., meeting of the International Congress of Entomology. She and several colleagues published a paper entitled “From the Students to the Students: Why YOU need to Attend ICE 2016.”
The Royal Entomological Society will publish her biography and photo in its Antenna magazine, on the society website, and in the Marsh Christian Trust Award brochure.
- Author: Kathy Keatley Garvey
Sarah Silverman, a doctoral candidate who is studying insect demography at UC Davis with major professor James R. Carey, won a second-place award in her category.
Boudinot, who studies classification and evolution of morphology, delivered a 10-minute oral presentation in the Systematics, Evolution and Biodiversity Section on "The Protopodal Theory of Genitalic Evolution in the Hexapoda (Arthropoda: Mandibulata: Pancrustacea)."
Boudinot completed his undergraduate work at the Evergreen State College, Olympia, Wash., and spent a year working as a research technician at the University of Utah before starting his graduate work in 2014 with advisor Phil Ward. He focuses his research on evolution and ecology, approached from the perspective of systematics. “I integrate several lines of inquiry to answer historical evolutionary questions, including morphological and molecular phylogenetics, paleontology, and traditional comparative morphology,” Boudinot related. “I specialize on the skeletomusculature system of the male genitalia of the Hexapoda and the classification of the Formicoidea.”
Silverman gave a 10-minute oral presentation in the Diptera-Mosquitoes category of the Medical, Urban, and Veterinary Entomology Section, on “Population as Cohort: Interpreting the Mortality Patterns of Wild-Caught Adult Mosquitoes of Unknown Ages.”
Silverman completed her bachelor's degree in environmental science at McGill University in Montreal. For her undergraduate thesis, she studied the phenology of wild Osmia bees. Her work at UC Davis is in the field of insect demography. “I specifically study insect lifespan in the wild, as well as the the age-structure of insect populations in the wild using an innovative methodological approach: the capture of live-insects in the wild which are then maintained and observed in the lab until death,” she said.
At its annual meetings, the ESA offers graduate students the opportunity to present their research and win coveted prizes. The first-place President's Prize recipient receives a one-year free membership in ESA, a $75 cash prize, and a certificate. The second-place winner receives a one-year free membership in ESA and a certificate.
The oral presentations are evaluated on scientific content (50 percent) and presentation (50 percent). For scientific content, judges score them on introduction and background with pertinent literature cited; objectives clearly stated and concise; materials and methods (study design) clear and concise; results and discussion clear, concise and accurate; and significance of results to field of study. Judges evaluation the presentation on organization, slides and delivery.
Boudinot's previous President's Prizes were for work on the male genitalia of ants, and for providing the first male-based identification material for the ant genera of the New World.
Ants are highly diverse, with over 13,000 known species, Boudinot said. "They are, however, but one stitch in the diversity of all insects, and we are entering a new era for the study of morphology in the 21st century."
The genitalia of male insects are fascinating, he said. "Both male and female insect genitalia are derived from the appendages of a pair of abdominal segments. Evidence from the skeletomusculature indicates that these structures are really legs of a crustacean ancestor that have been modified for numerous reproductive tasks--from copulation and insemination, to singing and silk-spinning."