Internationally acclaimed honey bee geneticist Robert E. Page Jr., a UC Davis alumnus and an emeritus administrator at UC Davis and Arizona State University, has been named the recipient of the 2023 C. W. Woodworth Award, the highest honor accorded by the Pacific Branch, Entomological Society of America (PBESA). 

“Dr. Page is a pioneering researcher in the field of evolutionary genetics and social behavior of honey bees, and a highly respected and quoted author, teacher and former administrator,” wrote nominator Steve Nadler, professor and chair of the UC Davis Department of Entomology and Nematology. 

Page is the 12th UC Davis recipient of the award, first presented in 1969. His mentor, and later colleague, Harry H. Laidlaw Jr. (for whom the UC Davis bee biology research facility is named), won the award in 1981.

The PBESA awards ceremony will take place at its meeting, April 2-5, in Seattle. The organization encompasses 11 Western states, and parts of Canada, Mexico and U.S. territories. 

“One of Dr. Page's most salient contributions to science was to construct the first genomic map of the honey bee, which sparked a variety of pioneering contributions not only to insect biology but to genetics at large,” Nadler related. “It was the first genetic map of any social insect. He was the first to demonstrate that a significant amount of observed behavioral variation among honey bee workers is due to genotypic variation. In the 1990s, he and his students and colleagues isolated, characterized and validated the complementary sex determination gene of the honey bee; considered the most important paper yet published about the genetics of Hymenoptera. The journal Cell featured their work on its cover. In subsequent studies, he and his team published further research into the regulation of honey bee foraging, defensive and alarm behavior.” 

A native of Bakersfield, Rob holds a bachelor's degree in entomology (1976), with a minor in chemistry, from San Jose State University. He obtained his doctorate in entomology (1980) from UC Davis; joined the UC Davis faculty in 1989; and chaired the Department of Entomology from 1999 to 2004. He gained UC Davis emeritus professor-chair status in 2004,  the year that Arizona State University (ASU) recruited him to be the founding director of its School of Life Sciences. Page organized three departments—biology, microbiology and botany, totaling more than 600 faculty, graduate students, postdoctoral fellows and staff--into one unified school. Using his UC Davis experiences, he established the school as a platform for discovery in the biomedical, genomic and evolutionary and environmental sciences; and established ASU's world-class Social Insect Research Group and Honey Bee Research Facility.

Page's career at ASU led to a series of top-level administrative roles: founding director, School of Life Sciences (2004-2010), vice provost and dean, College of Liberal Arts and Sciences (2011-2013) and university provost, 2014-2015.

Nadler praised Page's strategic vision, his leadership and his contributions to science.  He built two modern apicultural labs (in Ohio and Arizona), major legacies that are centers of honey bee research and training. The Social Insect Research Group (SIRG) at ASU is regarded as “the best in the world,” according to the late E. O. Wilson. ASU Professor Bert Hoelldobler, in an ASU news release, declared Dr. Page  as "the leading honey bee geneticist in the world. A number of now well-known scientists in the U.S. and Europe learned the ropes of sociogenetics in Rob's laboratory.”

Page is an Institute for Scientific Information (ISI) highly-cited author in plant and animal science. In all, he has published more than 250 research papers and articles, edited or authored five books, and is listed as a “highly-cited author” by the ISI Web of Knowledge, representing the top 1/2 of one percent of publishing scientists. Statistics include: H-index = 80, and 21,925 citations.  Since retiring from UC Davis, Dr. Page has published 65 research papers, 8 major reviews and 2 scholarly books, many using his UC Davis affiliation. He authored “The Spirit of the Hive: The Mechanisms of Social Evolution” (Harvard University Press, 2013) and the “Art of the Bee: Shaping the Environment from Landscapes to Societies” (Oxford University Press, 2020).

While at UC Davis, Page worked closely with his doctoral research mentor, Harry H. Laidlaw Jr., the father of honey bee genetics, and together they published many significant research papers and the landmark book, “Queen Rearing and Bee Breeding” (Wicwas Press, 1998). It is considered the most important resource book for honey bee genetics, breeding, and queen rearing.  Page is now in the process of updating it.

For 24 years, from 1989 to 2015, Page maintained a honey bee-breeding program, managed by bee breeder-geneticist Kim Fondrk. Their contributions include discovering a link between social behavior and maternal traits in bees. Their work was featured in a cover story in the journal Nature. In all, Nature featured his work on four covers from work mostly done at UC Davis.

A 2012 Fellow of the Entomological Society of America, Page has held national and international offices. He served as secretary, chair-elect, chair, subsection cb (apiculture and social insects) of ESA from 1986-1989; president of the North American Section, International Union for the Study of Social Insects, 1991; and a Council member, International Bee Research Association, 1995-2000.

Among his many honors:

• Fellow of the American Association for the Advancement of Science
• Awardee of the Alexander von Humboldt Senior Scientist Award (the Humboldt Prize - the highest honor given by the German government to foreign scientists)
• Foreign Member of the Brazilian Academy of Sciences
• Fellow of the American Academy of Arts and Sciences
• Elected to the Leopoldina - the German National Academy of Sciences (the longest continuing academy in the world)
• Fellow of the Wissenschaftskolleg zu Berlin
• Fellow of the Entomological Society of America
• Awardee of the Carl Friedrich von Siemens Fellowship
• Fellow of the California Academy of Sciences
• Fellow, Carl Friedrich von Siemens Foundation, Munich, Germany, September 2017-August
• Thomas and Nina Leigh Distinguished Alumni Award from UC Davis Department of Entomology and Nematology
• James Creasman Award of Excellence (ASU Alumni Association)
• UC Davis Distinguished Emeritus Professor, one awarded annually
• Distinguished Emeritus Award, College of Agricultural and Environmental Sciences, one awarded annually

In his letter of support, colleague and research collaborator James R. Carey, distinguished professor of entomology at UC Davis,  described Page as "one of the most gifted scientists, administrators, and teachers I have had the privilege to know in my 42 years in academia.”

Bee breeder-geneticist Susan Cobey of Washington State University, former manager of the Laidlaw facility, emphasized Page's importance to the bee breeding and beekeeping industry. Cobey, who has based her career on the Page-Laidlaw Closed Population Breeding, wrote that: “The beauty of this system is that it is practical and addresses the unique challenges of honey bee stock improvement. Queens mate in flight with numerous drones and selection is based upon complex behaviors at the colony level, influenced by the environmental. Hence, traditional animal breeding models do not apply well to honey bees.”

Nadler also noted that “Dr. Page was involved in genome mappings of bumble bees, parasitic wasps and two species of ants. His most recent work focuses on the genetic bases of individuality in honey bees; demonstrating genetic links between pollen and nectar collection, tactile and olfactory learning characteristics, and neuroendocrine function. This work provides the most detailed understanding to date of the molecular and genetic bases to task variation in a social insect colony.” 

Nadler added: "Not surprisingly, Dr. Page humbly considers his most far-reaching and important accomplishment, the success of his mentees, including at least 25 graduate students and postdocs who are now faculty members at leading research institutions around the world."

Charles William Woodworth (1865-1940), is considered the founder of both the UC Berkeley and UC Davis departments of entomology.  William Harry Lange Jr., (1912-2004) was the first UC Davis recipient of the Woodworth award (1978). Other recipients: Harry Laidlaw Jr., (1907-2003), 1981; Robert Washino, 1987; Thomas Leigh (1923-1993), 1991; Harry Kaya, 1998; Charles Summers, 1941-2021 (stationed at the UC Kearney Research and Extension throughout his career but a member of the UC Davis entomology faculty since 1992), 2009; Walter Leal, 2010; Frank Zalom, 2011; James R. Carey, 2014; Thomas Scott, 2015; and Lynn Kimsey, 2020.

The list of this year's PBESA recipients is posted here.

It was like coming home when acclaimed honey bee geneticist Robert E. Page Jr.,  stepped forward to accept the Exceptional Emeriti Faculty Award from Helene Dillard, dean of the UC Davis College of Agricultural and Environmental Sciences (CA&ES). 

The occasion: the college's Award of Distinction dinner and ceremony, held Nov. 3 in the Activities and Recreation Center Ballroom.

Page, considered by his peers as the world's leading honey bee geneticist, traces his "bee biology roots" to UC Davis. He drew a standing ovation.

Page received his doctorate in entomology in 1980 from UC Davis, studying with Harry H. Laidlaw Jr., and went on to join the UC Davis Department of Entomology faculty in 1989 and chair the department from 1999 to 2004. He then transitioned to emeritus and was recruited by Arizona State University (ASU)  to be the founding director of its School of Life Sciences. His career at ASU led to a series of top-level administrative roles: from founding director of the School of Life Sciences (2004-2010) to vice provost and dean, College of Liberal Arts and Sciences (2011-2013) and then to University Provost, 2014-2015.

"Rob is a pioneering researcher in the field of evolutionary genetics and social behavior of honey bees, and a highly respected and quoted author, teacher and former administrator,” wrote nominator Steve Nadler, professor and chair of the UC Davis Department of Entomology and Nematology.

“One of Dr. Page's most salient contributions to science was to construct the first genomic map of the honey bee, which sparked a variety of pioneering contributions not only to insect biology but to genetics at large,” Nadler pointed out. “It was the first genetic map of any social insect. He was the first to demonstrate that a significant amount of observed behavioral variation among honey bee workers is due to genotypic variation. In the 1990s, he and his students and colleagues isolated, characterized and validated the complementary sex determination gene of the honey bee; considered the most important paper yet published about the genetics of Hymenoptera. The journal Cell featured their work on its cover. In subsequent studies, he and his team published further research into the regulation of honey bee foraging, defensive and alarm behavior.”

While at UC Davis, Page worked closely with Laidlaw, "the father of honey bee genetics." Together they published many significant research papers and the landmark book, “Queen Rearing and Bee Breeding” (Wicwas Press, 1998), considered the most important resource book for honey bee genetics, breeding, and queen rearing.

For 24 years, from 1989 to 2015, Page maintained a UC Davis honey bee-breeding program, managed by bee breeder-geneticist Kim Fondrk. Their contributions include discovering a link between social behavior and maternal traits in bees. Their work was featured in a cover story in the journal Nature. In all, Nature featured his work on four covers from work mostly done at UC Davis. 

Since his retirement from UC Davis, Page has published 65 research papers, eight major reviews and two scholarly books, many using his UC Davis affiliation. He authored “The Spirit of the Hive: The Mechanisms of Social Evolution” (Harvard University Press, 2013) and the “Art of the Bee: Shaping the Environment from Landscapes to Societies” (Oxford University Press, 2020).

Now residing near Davis, Page continues to focus his research on honey bee behavior and population genetics, particularly the evolution of complex social behavior. His continuing academic activities bring credit to bee biology and UC Davis, Nadler said. “His large number of publications and citations continue to be an important component of the high national rating of our entomology department.”

To date, Page has published more than 250 research papers and articles, edited or authored five books, and is listed as a “highly-cited author” by the ISI Web of Knowledge, representing the top 1/2 of one percent of publishing scientists. 

Page continues to work closely with UC Davis professors and students, offering advice, helping them with grants, and editing manuscripts. A few years ago, he held an international workshop at the Laidlaw facility. He teaches courses (including “The Social Contract: from Rousseau to the Honey Bee,” and “The Song of the Queen: Thrilling Tales of Honey Bee Mating Behavior”) for the UC Davis Osher Lifelong Learning Institute (OLLI).

“Not surprisingly, Dr. Page humbly considers his most far-reaching and important accomplishment, the success of his mentees, including at least 25 graduate students and postdocs who are now faculty members at leading research institutions around the world,” Nadler wrote. “He also built two modern apicultural labs (in Ohio and Arizona), major legacies that are centers of honey bee research and training. He has trained many hundreds of beekeepers. His public service now extends to working as a Fellow with the California Academy of Sciences.”

Among Page's many honors:

• Fellow of the American Association for the Advancement of Science
• Awardee of the Alexander von Humboldt Senior Scientist Award (the Humboldt Prize - the highest honor given by the German government to foreign scientists)
• Foreign Member of the Brazilian Academy of Sciences
• Fellow of the American Academy of Arts and Sciences
• Elected to the Leopoldina, the German National Academy of Sciences (the longest continuing academy in the world)
• Fellow of the Wissenschaftskolleg zu Berlin
• Fellow of the Entomological Society of America
• Awardee of the Carl Friedrich von Siemens Fellowship
• Fellow of the California Academy of Sciences
• Fellow, Carl Friedrich von Siemens Foundation, Munich, Germany, September 2017-August 2018
• Thomas and Nina Leigh Distinguished Alumni Award from UC Davis Department of Entomology and Nematology
• James Creasman Award of Excellence (ASU Alumni Association)
• Regents Professor, Arizona State University
• UC Davis Distinguished Emeritus Professor

UC Davis-trained honey bee scientist Robert E. Page Jr., known as “the leading honey bee geneticist in the world,” has been named the recipient of the Exceptional Emeriti Faculty Award, from the UC Davis College of Agricultural and Environmental Sciences (CA&ES). 

He will be honored at the CA&ES Award of Distinction dinner and celebration awards ceremony on Thursday, Nov. 3 in the UC Davis Activities and Recreation Center Ballroom.

Page, emeritus professor and chair of the UC Davis Department of Entomology and an emeritus professor and administrator at Arizona State University, “is a pioneering researcher in the field of evolutionary genetics and social behavior of honey bees, and a highly respected and quoted author, teacher and former administrator,” wrote nominator Steve Nadler, professor and chair of the UC Davis Department of Entomology and Nematology.

“One of Dr. Page's most salient contributions to science was to construct the first genomic map of the honey bee, which sparked a variety of pioneering contributions not only to insect biology but to genetics at large,” Nadler pointed out. “It was the first genetic map of any social insect. He was the first to demonstrate that a significant amount of observed behavioral variation among honey bee workers is due to genotypic variation. In the 1990s, he and his students and colleagues isolated, characterized and validated the complementary sex determination gene of the honey bee; considered the most important paper yet published about the genetics of Hymenoptera. The journal Cell featured their work on its cover. In subsequent studies, he and his team published further research into the regulation of honey bee foraging, defensive and alarm behavior.”

A native of Bakersfield, Kern County, Rob holds a bachelor's degree in entomology (1976), with a minor in chemistry, from San Jose State University. He received his doctorate in entomology in 1980 from UC Davis, where he studied with his doctoral research mentor, Harry H. Laidlaw Jr.,  “the father of honey bee genetics” for whom the university's bee facility, the Harry H. Laidlaw Jr. Honey Bee Research Facility on Bee Biology Road, is named.

Page joined the UC Davis entomology faculty in 1989 and chaired the department from 1999 to 2004 when he transitioned to emeritus and was recruited by Arizona State University (ASU)  to be the founding director of its School of Life Sciences. His career at ASU led to a series of top-level administrative roles: from founding director of the School of Life Sciences (2004-2010) to vice provost and dean, College of Liberal Arts and Sciences (2011-2013) and then to University Provost, 2014-2015.

While at UC Davis, Page worked closely with Laidlaw. Together they published many significant research papers and the landmark book, “Queen Rearing and Bee Breeding” (Wicwas Press, 1998), considered the most important resource book for honey bee genetics, breeding, and queen rearing.

For 24 years, from 1989 to 2015, Page maintained a UC Davis honey bee-breeding program, managed by bee breeder-geneticist Kim Fondrk. Their contributions include discovering a link between social behavior and maternal traits in bees. Their work was featured in a cover story in the journal Nature. In all, Nature featured his work on four covers from work mostly done at UC Davis. 

Since his retirement from UC Davis, Page has published 65 research papers, eight major reviews and two scholarly books, many using his UC Davis affiliation. He authored “The Spirit of the Hive: The Mechanisms of Social Evolution” (Harvard University Press, 2013) and the “Art of the Bee: Shaping the Environment from Landscapes to Societies” (Oxford University Press, 2020).

Now residing near Davis, Page continues to focus his research on honey bee behavior and population genetics, particularly the evolution of complex social behavior. His continuing academic activities bring credit to bee biology and UC Davis, Nadler said. “His large number of publications and citations continue to be an important component of the high national rating of our entomology department.”

To date, Page has published more than 250 research papers and articles, edited or authored five books, and is listed as a “highly-cited author” by the ISI Web of Knowledge, representing the top 1/2 of one percent of publishing scientists.

Page continues to work closely with UC Davis professors and students, offering advice, helping them with grants, and editing manuscripts. A few years ago, he held an international workshop at the Laidlaw facility. He teaches courses (including “The Social Contract: from Rousseau to the Honey Bee,” and “The Song of the Queen: Thrilling Tales of Honey Bee Mating Behavior”) for the UC Davis Osher Lifelong Learning Institute (OLLI).

“Not surprisingly, Dr. Page humbly considers his most far-reaching and important accomplishment, the success of his mentees, including at least 25 graduate students and postdocs who are now faculty members at leading research institutions around the world,” Nadler wrote. “He also built two modern apicultural labs (in Ohio and Arizona), major legacies that are centers of honey bee research and training. He has trained many hundreds of beekeepers. His public service now extends to working as a Fellow with the California Academy of Sciences.”

Page's colleagues praise his strategic vision, his leadership and his contributions to science. ASU Professor Bert Hoelldobler described him as “the leading honey bee geneticist in the world. A number of now well-known scientists in the U.S. and Europe learned the ropes of sociogenetics in Rob's laboratory.”

“Rob is one of the most gifted scientists, administrators, and teachers I have had the privilege to know in my 42 years in academia,” said James R. Carey, a UC Davis distinguished professor of entomology.

Bee breeder-geneticist Susan Cobey of Washington State University, former manager of the Laidlaw facility, emphasized Page's importance to the bee breeding and beekeeping industry. She has based her entire career on the Page-Laidlaw Closed Population Breeding. “The beauty of this system is that it is practical and addresses the unique challenges of honey bee stock improvement,” she related. “Queens mate in flight with numerous drones and selection is based upon complex behaviors at the colony level, influenced by the environmental. Hence, traditional animal breeding models do not apply well to honey bees.”

When Page addressed the 2016 meeting of the International Congress of Entomology (ICE), he was introduced as “an engaging, energetic, humorous and highly intellectual speaker; this will be a memorable, entertaining and highly educational lecture by one of the most important entomologists of the 21st^t century. His background is in behavior and population genetics, and the focus of his current research is on the evolution of complex social behavior. Using the honey bee as a model, Dr. Page has dissected their complex foraging division of labor at all levels of biological organization, from gene networks to complex social interactions.”

Among Page's many honors:

• Fellow of the American Association for the Advancement of Science
• Awardee of the Alexander von Humboldt Senior Scientist Award (the Humboldt Prize - the highest honor given by the German government to foreign scientists)
• Foreign Member of the Brazilian Academy of Sciences
• Fellow of the American Academy of Arts and Sciences
• Elected to the Leopoldina, the German National Academy of Sciences (the longest continuing academy in the world)
• Fellow of the Wissenschaftskolleg zu Berlin
• Fellow of the Entomological Society of America
• Awardee of the Carl Friedrich von Siemens Fellowship
• Fellow of the California Academy of Sciences
• Fellow, Carl Friedrich von Siemens Foundation, Munich, Germany, September 2017-August 2018
• Thomas and Nina Leigh Distinguished Alumni Award from UC Davis Department of Entomology and Nematology
• James Creasman Award of Excellence (ASU Alumni Association)
• Regents Professor, Arizona State University
• UC Davis Distinguished Emeritus Professor

Two geneticists affiliated with the UC Davis Department of Entomology and Nematology are each scheduled to teach a 2022 virtual course as part of the Osher Lifelong Learning Institute (OLLI),  which offers educational programs for Davis area seniors, ages 50 and over.

Honey bee geneticist and UC Davis alumnus Robert E. Page Jr., will teach a course on "The Social Contract: from Rousseau to the Honey Bee" on Monday, Feb. 7 from 2 to 4 p.m.  He is an emeritus professor and former chair of the UC Davis Department of Entomology and Nematology and a university provost emeritus and a regents' professor emeritus at Arizona State University.  A UC Davis alumnus and the 2019 recipient of the UC Davis Distinguished Emeritus Award, administered by the UC Davis Emeriti Association, Page has published more than 230 research papers and three books on honey bee genetics and behavior. Course specifics: 213SNR388 $25 Zoom.

Molecular geneticist and physiologist Joanna Chiu, professor and vice chair of the UC Davis Department of Entomology and Nematology and a Chancellor's Fellow, will cover "The Circadian Clock and Chronomedicine" on Wednesday, March 23 from 10 a.m. to noon. Chiu, who holds a doctorate in molecular genetics from New York University, received postdoctoral training at the Center for Advanced Biotechnology and Medicine at Rutgers University.  Course specifics: 213SNR383 $25 Zoom.

Social Contract: from Rousseau to the Honey Bee Animals 
"Animals that live together in a society, like the social insects, have a tacit agreement, a social contract, that guarantees that their interests are protected in exchange for their social cooperation," Page says. "Like social contracts that bind human societies, this contract isn't written on paper, it is implied, though in humans its enforcement is expressed in explicit written laws and national constitutions. The social contract of insects has been written by natural selection in their DNA over thousands of generations. In this class we will explore elements of the social contract of honey bee societies, its origins and 'laws for enforcement,' and the social services honey bees get as a consequence of being members of a society. Specifically, we will explore their systems of national defense, internal police, public works, public health and border control. Without these features, their society would fail, as would ours. Discover what we can learn about ourselves from studying social insects."

The Circadian Clock and Chronomedicine
"The circadian clock is an internal body clock that controls all aspects of physiology and behavior in every single one of us. It interprets environmental and metabolic time cues to ensure our body performs at its best," Chiu says. "A sequel to the OLLI course 'What Time is it and Why Does it Matter?,' this course will revisit the very important, yet infrequently discussed physiological machinery that controls sleep- wake cycles, hormone production and immune system. Because disruptions in the circadian clock have been associated with a range of human diseases, we will highlight life-style choices and work schedules that cause clock disruptions. Finally, we will discuss new strategies in the medical field that leverage circadian biology concepts to increase the efficacy of medical treatments."

OLLI is headquartered at 463 California Ave. Davis, CA 95616.  Members of the UC Davis Retirees' Association and the UC Davis Emeriti Association are offered a complimentary course credit equivalent to their paid OLLI membership fee.

The annual OLLI membership is $60 for the academic year (Oct. 4, 2021- June 30, 2022). More information on OLLI, which is part of the UC Davis Continuing and Professional Education, is available online or contact (530) 752-9695 or olli@ucdavis.edu