- Author: Kathy Keatley Garvey
Wilson, recognized as one of the world's most influential scientists, was known as “The Ant Man,” "The Father of Sociobiology," "The Father of Diversity" and “The Modern-Day Darwin," for his pioneering and trailblazing work that drew global admiration and won scores of scientific awards.
But among his peers, colleagues and mentees, he was known as "Ed."
Wilson's work, On Human Nature, won the Pulitzer Prize in 1979. He won a second Pulitzer in 1991 with The Ants, co-authored with colleague Bert Hölldobler. In 1990, the Royal Swedish Academy of Sciences awarded Wilson the Crafoord Prize in biosciences, the highest scientific award in the field. In 1996, Time magazine named him one of America's 25 most influential people. In 1977 President Jimmy Carter awarded him the National Medal of Science for his contributions toward the advancement of knowledge in biology.
Wilson, according to reports, always considered himself an Alabaman who went to Harvard, rather than a Harvard professor born in Alabama. Born June 10, 1929 in Birmingham, Ed graduated from the University of Alabama in 1949 with two degrees in biology, and received his doctorate in biology from Harvard in 1955. He joined the Harvard faculty in 1956. Although officially retiring in 1996, he remained active as an emeritus professor and honorary curator until his death.
Some tributes from UC Davis faculty and students:
“E. O. Wilson was a towering figure in the study of social insects, in evolutionary biology, and in conservation biology,” said fellow ant specialist and professor Phil Ward of the UC Davis Department of Entomology and Nematology who organized the 2007 E.O. Wilson Festschrift (a collection of writings published in honor of a scholar). “He made important contributions in all of these areas, but his specialty was the study of ants, those ‘little creatures that run the world.' Wilson's book, The Insect Societies (1971), introduced its readers to the fascinating world of ants and other social insects, using language that was both engaging and accessible, yet highly informative.”
“This was followed two decades later by the equally magisterial The Ants, co-authored with Bert Hölldobler,” Ward noted. “These landmark contributions inspired many budding biologists, myself included, to devote ourselves to the study of ants and other social organisms. Equally important, Wilson argued passionately and compellingly for the conservation of biological diversity in a dwindling natural world. He once said that 'destroying rainforest for economic gain is like burning a Renaissance painting to cook a meal.' Let us honor his legacy by heeding this message!”
UC Davis doctoral alumnus Brendon Boudinot of the Phil Ward lab and now a postdoctoral researcher in Germany at the Institute of Zoology and Evolutionary Research, Friedrich Schiller University Jena, says the ant world is reeling with Wilson's passing. “A big chunk of my dissertation was dedicated to testing his hypotheses for the origin and early evolution of ants!”
Boudinot met Wilson when he was visiting the Ant Room at the Harvard Museum of Comparative Zoology in 2013. “The work I was doing was the foundation for my studies on ant males, and I was near the end of the trip, at one of the many microscopes by the window facing the yard,” he said. “Ed surprised me by coming right up to my shoulder at the scope; he asked me what I was working on. I am a bit abashed to say that I couldn't say anything because my mind went blank! Stef Cover, the pins-and-points curator told Ed what I was doing, and for the life of me I will always remember what Wilson said. He was happy that I chose to work on male ants, when this sex has been actively ignored by researchers over the past centuries, and that he himself was more apt to squash one at a light trap than to collect one. I hope that my keys and diagnoses have helped people appreciate male ants.”
“I am so thankful that I met him,” Boudinot said, “and that I was able to work with so many people in his sphere. The ant world is reeling, as he was a gentle giant of myrmecology, and of course biology writ large.”
Doctoral candidate Jill Oberski of the Phil Ward lab met “The Ant Man” in 2019. “I got to meet E. O. Wilson when I traveled to the Harvard Museum of Comparative Zoology in 2019. In addition to myself, there were several other researchers visiting the “Ant Room,” which houses a huge number of type specimens.”
“He asked me about my research on Dorymyrmex taxonomy and biogeography, although as a second-year PhD student I didn't have much to report yet. He was genuinely interested in my work and excited that I was working to resolve Dorymyrmex--which has always been a taxonomic headache. He also told me he recalled watching ants forming cone-shaped nests as a child in Alabama, which could only have been Dorymyrmex. He was exceedingly kind and encouraging.
“Finally, the Ant Room staff and visitors ate lunch together in Ed's office—lobster sandwiches, diet Dr. Pepper, and coffee, as is customary.”
Lynn Kimsey, director of the Bohart Museum of Entomology and a UC Davis distinguished professor of entomology, remembers working near his office when she served as a visiting professor/lecturer (1987 to 1989) at Harvard's Museum of Comparative Zoology, before joining the UC Davis faculty in 1989.
“His office was just down the hall from mine when I taught at Harvard. Here he was, one of the most famous biologists of his generation and I would see him sit down on the sidewalk to show a little kid the ants there. Also, saw him in the sitting on a bench Burlington Mall while his wife shopped, writing on a yellow pad of paper. Totally focused on what he was writing with shopping pandemonium all around. He was brilliant, humble and engaging.”
UC Davis doctoral alumnus Fran Keller, a professor at Folsom Lake College, and a research scientist at the Bohart Museum of Entomology, helped honor his work at a special symposium hosted at the 2005 Entomological Society of America meeting in Fort Lauderdale, Fla.
"Our department of entomology helped fund my trip to Harvard,” she recalled, “and he agreed to meet me over the course of two days in May 2005. The ESA symposium took place in mid-December. I recorded our interview on a cassette tape,” she said, adding she hopes to publish it in a journal.
Wilson responded: “Everyone has an animal that reaches them or that they connect with at some level, even though you were born an entomologist, perhaps yours is the rhino.”
“After my interview with Ed, I bought the book in the MCZ, The Rarest of the Rare: Stories Behind the Treasues at the Harvard Museum of Natural History. In that book, it highlights the extinct and rare species held in the MCZ collections. One of those specimens is the last Xerces butterfly, which was caught by Harry Lange (UC Davis emeritus professor of entomology). Harry's quote in that book, ‘I didn't know it was the last one, I thought there would be more' and then my time eating lunch and then wandering the MCZ collection and chatting with Ed inspired me to create the Xerces t-shirt for the Bohart Museum of Entomology.”
One of Keller's mentors, Tom Schoener, studied with Wilson. “I worked on plant ecology and island biogeography for my undergrad research (Sacramento City College)," she said, "and continued that for awhile in grad school (UC Davis). Ed Wilson was one of the founders of island biogeography.”
As a undergraduate at Sacramento City College, Keller was part of a field trip to hear Wilson speak at his 2002 book tour on The Future of Life.
UC Davis doctoral alumnus Alex Wild, an evolutionary biologist, science photographer and curator of entomology at the University of Texas, Austin, wrote this on his Twitter account, @Myrmecos, which has more than 30,000 followers: “Ed Wilson was one of my science heroes. Over the years I came to admire two things in particular. One was his ability to craft technical books so compelling as to launch scores of scientific careers in their wake.”
“The other thing is how Ed Wilson handled professional disagreement," Wild tweeted. "And he had a lot of those, because Wilson was frequently wrong. About a great range of topics. For a guy known for ant research, his interpretation of ant origins was just… silly.”
"But he continued to support, both financially and professionally, the young upstarts who, over and over, proved him wrong. That's a rare trait for a field as ego-driven as evolutionary biology.”
When a follower asked: “Can you explain to a non-biologist bug enthusiast why his interpretation of ant origins was silly?”, Wild replied: “His arrangement of the ant subfamilies, based on subjective hunches of evolutionary relationships rather than data, bore no resemblance at all to the well-supported relationships from subsequent data-based studies, like https://www.pnas.org/content/103/48/18172. (This 2006 research article, "Evaluating Alternative Hypotheses for the Early Evolution and Diversification of Ants," is co-authored by Seán G. Brady, Ted R. Schultz, Brian L. Fisher, and Philip S. Ward and edited by Bert Hölldobler, University of Würzburg, Würzburg, Germany)
“There are no words that adequately describe E.O. Wilson's courage in challenging dogma, his energy in documenting and sharing the wonders of our planet, or his extraordinary creativity," said Diane Ullman, UC Davis professor of entomology and former chair of the Department of Entomology. "So much of his writing touched me deeply, from his writing about the continuum between art and science (Consilience), to his collaboration with Bert Hölldobler, addressing the incredible biology and behavior of social insects (Superorganism). He wrote a wonderful, 'coming of age' novel, seemingly much inspired by his own youth (Anthill)."
"The first time I was able to hear him speak in person was in 1996 at the International Congress of Entomology in Florence, Italy, where he was the plenary speaker opening the meeting. He spoke passionately about loss of biodiversity, and was sounding the alarm on impact of humans and climate change on the planet. He continued to lead this charge up to the very end, never giving up on proposing potential solutions on a global scale. He was witty and with use of metaphor made us see so very many things."
Caleb Johnson, who teaches writing at Appalachian State University, Boone, N.C., described E.O. Wilson as “the world's forestmost authority on biodiversity” in an April 21, 2020 article in The Bitter Southerner. He referred to him as “A world-renowned scientific thinker whose vision for stopping this unprecedented environmental hemorrhaging is based on more than seven decades of careful witness, writing, and work in ecology and conservation."
Johnson wrote that Wilson lost his right eye in a fishing accident in the summer of 1936 near Paradise Beach, Fla., but he never let that stop his goals.
"In the 1940s, E.O. Wilson was an Alabama teenager who wandered the bottomland around Mobile and studied its creatures. He never stopped and became the world's foremost authority on biodiversity. He's 90 now, but still working, because he knows there's a way to undo the damage we've done to Mother Earth."
- Author: Kathy Keatley Garvey
- Asian giant hornets (aka "murder hornets"): 1 to 2 p.m., Thursday, Feb. 18, by Professor Lynn Kimsey, director of the Bohart Museum of Entomology and professor, UC Davis Department of Entomology and Nematology
- Ants: 11 a.m., to noon, Saturday, Feb. 20, by Professor Phil Ward, UC Davis Department of Entomology and Nematology
- Bees: 12:15 to 12:45, Tuesday, Feb. 23, by Christine Casey, manager of the Häagen-Dazs Honey Bee Haven
- Botanical Conservatory (in Spanish): 1 to 2 p.m., Wednesday, Feb. 23, by collections manager Ernesto Sandoval (postponed)
To obtain the Zoom links, click here.
About UC Davis Biodiversity Museum Month
The 10th annual UC Davis Biodiversity Museum Month program is all virtual this year via live talks and demonstrations, and pre-recorded presentations. It's being celebrated throughout the month of February. The science-based event traditionally occurs on only one day--the Saturday of Presidents' Weekend, when families and friends gather on campus to learn first-hand about the UC Davis museums and collections. The 2020 event drew more than 4000 to the campus.
This year's biodiversity event is featuring 12 museums or collections:
- Anthropology Museum
- Arboretum and Public Garden
- Bohart Museum of Entomology
- Botanical Conservatory
- California Raptor Center
- Center for Plant Diversity
- Häagen-Dazs Honey Bee Haven
- Nematode Collection
- Marine Invertebrate Collection
- Museum of Wildlife and Fish Biology
- Paleontology Collection
- Phaff Yeast Culture Collection
For more information and the schedule, access these two formats on the UC Davis Biodiversity program website: (1) live talks and demonstrations and (2) pre-recorded talks and activities. Information on the biodiversity museum events also appear on social media, including Facebook, Instagram, and Twitter, @BioDivDay.
If you'd like to donate to the UC Davis Diversity Museum Program in its crowdfunding efforts--this year is the 10th annual--click here. To donate to the Häagen-Dazs Honey Bee Haven, click here. To donate to the California Master Beekeeper Program, directed by Extension apiculturist Elina Lastro Niño of the UC Davis Department of Entomology and Nematology, click here. Niño also serves as the director of the haven. Crowdfunding will continue through the month of February.
- Author: Kathy Keatley Garvey
Boudinot, who studied with major professor Phil Ward of the UC Davis Department of Entomology and Nematology, is the second UC Davis-affiliated entomologist to receive the honor in its 28-year history. Jessica Gillung of the Lynn Kimsey lab, Bohart Museum of Entomology, won the award in 2019.
Snodgrass (1875-1962), a leader in insect morphology, is known for his 76 scientific articles and six books, including Insects, Their Ways and Means of Living (1930) and the book considered to be his crowning achievement, the Principles of Insect Morphology (1935).
Boudinot has them all. “Principles of Insect Morphology and the Anatomy of the Honey Bee were the foundation of my studies,” he said. “I have both, plus his Textbook of Arthropod Morphology and Insects, Their Ways and Means of Living on my desk in the lab.”
The Snodgrass Award, which includes a certificate and cash prize, recognizes outstanding research by a PhD student who has completed a research thesis or dissertation in arthropod morphology, systematics, taxonomy, or evolution. Nominees are scored on honors, awards, achievements and recognition; recommendations of professors and advisors; grantsmanship, publications, creativity and innovation of thesis or dissertation; and contribution to morphology.
Boudinot's dissertation: “Systematic and Evolutionary Morphology: Case Studies on Formicidae, Mesozoic Aculeata, and Hexapodan Genitalia.”
He earlier received the prestigious Alexander von Humboldt Research Fellowship to do research on evolutionary and comparative anatomy in Jena, Germany. He will locate to Germany in early 2021 for the two-year fellowship, after completing intensive language studies.
'I Am a Morphologist Because of Robert Evans Snodgrass'
“I am a morphologist because of Robert Evans Snodgrass,” Boudinot wrote in his application. “Although I have had pressure from my earliest undergraduate to become a molecular systematist, it was my chance encounter with Snodgrass's Principles of Insect Morphology late one night in the college library that set the course of my career. I had struggled for years at that point to understand the biodiversity of insects and to untangle the deep morass of arcane terminology, but my vision was illuminated by the conceptual clarity, linguistic simplicity, and exceptional engravings of the Principles. This work continues to hold special dominion over my thinking, as it is through the principle of mechanical function for explaining comparative anatomical observations that I have come to my present understanding of systematic and evolutionary morphology.”
Boudinot wrote that his “career objective, in brief, is to resolve the morphological evolution of insects through the synthesis traditional morphology, as represented by Snodgrass, with recent trends in digital anatomy and bioinformatics. I envision a future wherein students rely not on Borror & Delong, a holdover from Comstock's 19thCentury manual, but rather learn about insect structure, function, classification, and evolution through manipulation of three- and four-dimensional digital anatomical models, substantiated via manual curation and dissection. I want students to see for themselves what I have understood through the study of Snodgrass's work, balanced by contemporary research: The origin of the Arthropoda and morphological transformation through geological time to the resplendent, and endangered, diversity of today.
“In sum, my identity as an entomologist, and as a scientist more broadly, is due to the insights on the language and phenomenology of morphological evolution I gained from the oeuvre of Snodgrass. Without these works, I would probably still be a botanist.”
Boudinot's research interests include the origin and evolution of complex phenotypic systems from the perspective of phylogenetic systematics, including molecular and paleontological evidence. Specializing on morphological identity and evolutionary transformation, the skeletomuscular system of Arthropoda, with emphasis on the male genitalia of Hexapoda and systematics of the Hymenoptera, particularly the Formicidae.
John Henry Comstock Award
Highly honored for his academics, leadership, public service activities, professional activities and publications while at UC Davis, Boudinot received the 2019 John Henry Comstock Award, the top graduate student award given by ESA's Pacific Branch. The branch encompasses 11 Western states, U.S. territories, and parts of Canada and Mexico.
In the Comstock award application, Steve Nadler, professor and chair of the Department of Entomology and Nematology, described Boudinot as “A highly respected scientist, teacher and leader with a keen intellect, unbridled enthusiasm, and an incredible penchant for public service.” Boudinot maintained a 4.00 grade point average and published 18 peer-reviewed publications on insect systematics, some landmark or groundbreaking work.
His most recent publications: one on Cretaceous Strepsiptera in Cladistics and the other on the iron maiden ants in Myrmecological News ("Two New Iron Maiden Ants from Burmese Amber (Hymenoptera:
Formicidae: †Zigrasimeciini").
Boudinot received multiple “President's Prize” awards for his research presentations at national ESA meetings. He organized the ESA symposium, “Evolutionary and Phylogenetic Morphology,” at the 2018 meeting in Vancouver, B.C. , and delivered a presentation on “Male Ants: Past, Present and Prospects” at the 2016 International Congress of Entomology meeting in Orlando, Fla.
Boudinot served on—and anchored—three of the UC Davis Linnaean Games teams that won national or international ESA championships. The Linnaean Games, now known as the Entomology Games, are a lively question-and-answer, college bowl-style competitions on entomological facts played between university-sponsored student teams.
Brendon served as president of the UC Davis Entomology Graduate Student Association from 2006 to 2019, and co-chaired the department's UC Davis Picnic Day celebration (with forensic entomologist Robert Kimsey) for three years.
Before enrolling in graduate school program at UC Davis in 2014, Brendon received his bachelor's degree in entomology in 2012 from The Evergreen State College, Olympia, Wash. Professor John T. Longino served as his mentor.
- Author: Kathy Keatley Garvey
Boudinot presented a talk in Germany at the 9th Dresden Meeting on Insect Phylogeny, hosted by the Senckenberg Institute. In his presentation, "Skeletomuscular Evolution of Male Insect Genitalia, with Emphasis on the Endopterygota," he explained the second chapter of his thesis to an international audience of entomologists, spanning phylogeneticists to anatomists.
At the XXIV Simpósio de Mirmecologia in Belo Horizonte, Minas Gerais, Brazil, Boudinot will present preliminary results from his third and final thesis chapter in a symposium titled "Morphology Reloaded: Conquering the Third Dimension of Ant Evolution and Biogeography." His talk is titled "Altered States: Aculeate Anatomy and the Early Evolution of the Formicoidea."
"While this symposium is focused on the application of scanning technologies, such as X-ray computed microtomography, I will be speaking about the evolution of ant morphology through evolutionary time," Boudinot related. He will be presenting results from combined phylogenetic analysis of genome-scale DNA sequence data and a massive dataset of 576 morphological characters scored for 573 species.
"One of the novel aspects of my study is how many fossils I have included (more than 300)," Boudinot said. "The objective of this particular work is to estimate the geological age in which the ants originated, as well as the physical features of the ancestral ant, which I will then use to trace evolutionary and paleoecological patterns to the modern fauna."
Boudinot received the prestigious John Henry Comstock Award in April at the Pacific Branch, Entomological Society of America (PBESA) meeting in San Diego. This is the highest graduate student award given by PBESA, which encompasses 11 Western states, U.S. territories, and parts of Canada and Mexico. Its parent organization, the Entomological Society of America (ESA), will honor the six Comstock award winners--one from each branch--at its Nov. 17-20 meeting in St. Louis.
Boudinot, who was praised for his academic record, leadership, public service activities, participation in professional activities, and his publications, is the sixth UC Davis recipient of the Comstock award:
2019: Brendon Boudinot (major professor Phil Ward)
2015: Mohammad-Amir Aghaee (major professor the late Larry Godfrey)
2014: Kelly Hamby (major professor Frank Zalom)
2013: Matan Shelomi (major professor Lynn Kimsey)
2008: Christopher Barker (major professor William Reisen)
1983: Elaine Backus (major professor the late Donald McLean)
Active in PBESA and ESA, Boudinot received multiple “President's Prize” awards for his research presentations at national ESA meetings. He organized the ESA symposium, “Evolutionary and Phylogenetic Morphology,” at the 2018 meeting in Vancouver, B.C. , and delivered a presentation on “Male Ants: Past, Present and Prospects” at the 2016 International Congress of Entomology meeting in Orlando, Fla.
Boudinot served on—and anchored—three of the UC Davis Linnaean Games teams that won national or international ESA championships. The Linnaean Games are a lively question-and-answer, college bowl-style competition on entomological facts played between university-sponsored student teams.
Boudinot has served as president of the UC Davis Entomology Graduate Student Association since 2006, and is active in the campuswide UC Davis Picnic Day; he has co-chaired the department's Picnic Day Committee since 2017.
- Author: Kathy Keatley Garvey
The award, given by ESA's Systematics, Evolution and Biodiversity Section, recognizes outstanding research by a doctoral student who has completed a research thesis or dissertation in arthropod morphology, systematics, taxonomy, or evolution. The prize is a $500 cash award and an invited talk at ESA. She'll speak on "Unraveling the Evolution of Spider flies (Diptera, Acroceridae): Progress and Possibilities" at the ESA's annual meeting, set Nov. 17-20 in St. Louis, Mo.
Gillung is the first UC Davis student to receive the award, which was first presented in 1992. Her dissertation, “Systematics and Phylogenomics of Spider Flies (Diptera, Acroceridae),” focused on the evolution, conservation, biology, and taxonomy of spider flies, a group of spider natural enemies.
Awarded her doctorate in entomology in December 2018, Gilllung studied with major professor Lynn Kimsey, director of the Bohart Museum of Entomology and professor of entomology, UC Davis Department of Entomology and Nematology. She also studied with mentor Shaun Winterton, insect biosystematist, California Department of Food and Agriculture, and collaborated with ant specialist and taxonomist Phil Ward, professor, UC Davis Department of Entomology and Nematology.
Gillung's dissertation, involving genomics, phylogenetics, systematics, and comparative analyses, “has increased our understanding of the biological patterns and processes that have shaped our planet's biodiversity,” Kimsey wrote in her letter of nomination. Her taxonomic work included identification keys and morphology-based diagnoses of species using modern techniques of cybertaxonomy—the application of the internet, digital technologies, and computer resources to increase and speed up the discovery and cataloging of new species, Kimsey wrote in her letter of nomination.
“Using cybertaxonomic tools, Jessica described 25 new spider fly species herself, and in collaboration with fellow entomologists, three fossil species from Baltic amber, described in her first dissertation chapter," Kimsey wrote. "Cybertaxonomy is a powerful tool that allows researchers and citizen scientists to collaborate in real time and across great distances to increase the speed and efficiency of biodiversity discovery.”
Kimsey noted that “Jessica unraveled the functional and ecological implications of key morphological traits, as well as their distribution across the Tree of Life," and "established new homologies for the wing venation of spider flies. She conducted detailed and assiduous dissections of male reproductive structures (i.e., genitalia) to understand homologies, demonstrating that morphological traits are dynamically evolving systems useful for both classification and inference of evolutionary history.”
While at UC Davis, Gillung drew more $120,000 in grants and awards for her multifaceted research on genomics, bioinformatics, phylogenetics, plant-pollinator interactions, and biodiversity discovery. She compiled a near straight-A academic record, published 11 refereed publications in top journals, and engaged in public service and outreach programs that reached more than 20,000 people at UC Davis-based events.
Gillung was a key member of the UC Davis Linnaean Games Team that won the ESA national championship in 2015. The Linnaean Games are lively question-and-answer, college bowl-style competitions on entomological facts played between university-sponsored student teams.
The UC Davis-trained entomologist earlier received
- The 2019 Marsh Award for Early Career Entomologist, sponsored by the Royal Entomological Society. That involved a $1624 cash award and an invitation to the society meeting, Aug. 20-22 at the London School of Hygiene and Tropical Medicine.
- The 2019 Early Career Award from the Pacific Branch, Entomological Society of America (PBESA). PBESA encompasses 11 western states, U.S. territories and parts of Canada and Mexico.
- The 2018 Student Leadership Award from PBESA
- The “Best Student Presentation Award” at the ninth annual International Congress of Dipterology, held in 2018 in Windhoek, Namibia.
At Cornell, Gillung is researching Apoidea (stinging wasps and bees) phylogenomics, evolution and diversification in the Brian Danforth lab.