- Author: Ben Faber
There are 4,000 species of earthworms grouped into five families and distributed all over the world. Some grow uo to 3 feet long, while others are only a few tenths of inches. We call them nightcrawlers, field worms, manure worms, red worms and some people call them little diggers.
In California, we have some native species of earthworms, but in many cases non-native introduced species have come to dominate. The predominant native species belong to the Argilophilus and Diplocardia while many of the non-native are of European in origin in the Lumbricidae family. Many of these non-natives were probably introduced by settlers bringing plants from home, which had soil containing the worms. A survey of California earthworms by the US Forest Service can be found at:
https://www.fs.fed.us/psw/publications/documents/psw_gtr142/psw_gtr142.pdf
This is a wonderful description of earthworm biology and their occurrence in the landscape.
When digging in citrus orchards, it is common to find earthworms in the wetted mulch under tree canopies. Many of our citrus orchards were initially established by “balled and burlap” nursery trees that brought worms along with the soil. In the case of many avocado orchards, on the other hand, it can be rare to find earthworms in orchards. Most avocado orchards have been established since the 1970s when potting mixes and plastic liners were the standard practice and worms were not part of the planting media. Even though there is a thick leaf mulch in avocado orchards, the worms have not been introduced, and it is rare to find them.
Numerous investigators have pointed out the beneficial effects of earthworms on soil properties. One of the first of these observers was Charles Darwin who published Earthworms and Vegetable Mould in 1881. He remarked on the great quantity of soil the worms can move in a year. He estimated that the earthworms in some of his pastures could form a new layer of soil 7 inches thick in thirty years, or that they brought up about 20 tons of soil per acre, enough to form a layer 0.2-inch-deep each year.
Earthworms, where they flourish, are important agents in mixing the dead surface litter with the main body of the soil. They drag the leaves and other litter down into their burrows where soil microorganisms also begin digesting the material. Some earthworms can burrow as deeply as 5 to 6 feet, but most concentrate in the top 6 to 8 inches of soil.
The worm subsists on organic matter such as leaves and dead roots near the soil surface. The earthworm ingests soil particles along with the organic matter and grinds up the organic matter in a gizzard just as a chicken does. This is excreted in what we call worm casts. The castings differ chemically from the rest of the soil, as they are richer in nitrogen, potassium and other mineral constituents.
Castings are a natural by-product of worms. When added to normal soils in gardens or lawns, they provide the same kinds of benefits as other bulky organic fertilizers. Castings today are not commonly used as fertilizer by commercial plant growers because of their cost relative to other fertilizers. However, castings are used by some organic growers and are sold commercially as a soil amendment or planting medium for ornamental plants grown in pots.
The physical soil churning process also has several important effects:
-Organic residues are more rapidly degraded with the release of elements such as nitrogen, sulfur and other nutrients.
-Some of the inorganic soil minerals tend to be solubilized by the digestive process.
-Extensive burrowing improves soil aeration.
-Burrowing can improve water penetration into soils
-The earthworm carries surface nutrients from the soil surface and imports them into the root zone of the plant.
Although earthworms are considered beneficial to soil productivity, few valid studies have been made to determine whether their presence will significantly improve plant growth. This may seem odd since many of us have learned from childhood that worms are good. It is something like the chicken and the egg analogy. The conditions that are conducive to earthworms are also ideal for plants. Both plants and worms need temperatures between 60 and 100 degrees F for good growth; both need water, but not too much or little; they both require oxygen for respiration; and they do not like soils that are too acid or basic or too salty. By correcting soil conditions that are unfavorable for one will also improve the outlook for the other. The earthworm is a natural component of the soil population. If the soil is properly managed this natural population will thrive. In this sense, the presence or absence or earthworms can be an indicator of the "fertility" of one's soil.
- Author: Ben Faber
This is a reminder of the complexity of huanglongbing and the bacterial infection it causes. This abstract is from the HLB Conference in Florida last fall.
4.a.5 Symptom variations and molecular markers that illustrate the HLB complexity
Yongping Duan, Marco Pitino, and Cheryl Armstrong
USHRL-ARS-USDA, Fort Pierce, FL 34945, USA
Huanglongbing (HLB) is a devastating bacterial disease of citrus worldwide due to its intracellular and systemic infection. Various HLB symptoms are observed on different species/varieties of citrus plants: from yellow shoots to blotchy mottle on the leaves, from vein yellowing/vein corky to mosaic/green islands similar to zinc deficiency on the leaves, from whitish discoloration to stunted green leaves, etc. These variations of symptoms, which result from a combination of biotic and abiotic stresses, are not only present on individual plants from a variety but also exist on individual branches of an infected plant. Our results indicated that the adaptation of the bacterial populations, such as the dynamics of ‘Candidatus Liberibacter asiaticus' (Las), plays an important role in the induction of various symptoms and that Las mutations as well as the number and recombination events of Las prophages/phages affect this phenomenon. In addition, the selection of the host plants (resistance/tolerance) for the bacterial populations is also critical for symptom expression during disease progression. Based on severity, we divided HLB symptoms into four grades. It is worth noting that the grades of HLB symptom severity show a positive correlation with our newly identified biomarkers from host plants, and that gene expression profiling of different grades of infected leaves rationalized the differentiation based on the dynamics of these biomarkers. Because of these findings, we propose new approaches that allow for rapid selection of variant citrus plants, including bud sports with greater HLB resistance/tolerance.
Non-Technical Summary: Various symptoms of citrus huanglongbing display in different species/varieties of infected citrus plants. These variations of symptoms are not only present on individual plants from a variety, but also exist on individual branches of an infected plant. We have identified some molecular markers from the citrus plants and Las pathogen that illustrate the HLB complexity. Therefore, we propose new approaches that allow for rapid selection of variant citrus plants, including bud sports with greater HLB resistance/tolerance.
http://www.icc2016.com/images/icc2016/downloads/Abstract_Book_ICC_2016.pdf
- Author: Ben Faber
At the recent HLB Conference in Florida a paper was given that reinforces the need for appropriate soil and water pH to maximize root density and tree health. The industry there is dominated by a range of rootstocks and by Valencia-like varieties. Jim Graham and colleagues have shown that pH contributes to orchard health in their HLB situation. This should be a reminder for California growers for general tree health. Florida soils tend to be more coarse than soils found in many California orchards. It's much harder to change soil pH with acidified irrigation water with heavier textured soils.
4.b.1 Soil and water acidification sustain root density of huanglongbing-infected trees in Florida
Jim GRAHAM, Kayla GERBERICH, Diane BRIGHT, Evan JOHNSON
University of Florida, Citrus Research and Education Center, Lake Alfred, Florida, USA
Abstract: Early symptoms of HLB include fibrous root loss and leaf blotchy mottle, followed by premature fruit and leaf drop, and yield decline. As a consequence of initial bacterial infection of fibrous roots, a 30-50% reduction in fibrous root density and elevated soil Phytophthora populations were detected in field surveys. Continued sampling of Hamlin and Valencia orange trees on Swingle citrumelo rootstock in different stages of HLB decline revealed that root loss occurs in two stages. The second phase of root loss (70-80%) begins at the early stage of tree canopy thinning resulting from leaf drop and branch dieback. A more extensive survey of HLB-affected groves indicated that greater decline in fibrous root health and expression of HLB symptoms is observed where irrigation water is high in bicarbonates (> 100 ppm) and/or soil pH > 6.5. HLB symptom expression of trees on different rootstocks follows the known intolerance to bicarbonate (Swingle citrumelo > Carrizo citrange > sour orange > Cleopatra mandarin). Acidification of irrigation water in central ridge and south central flatwoods Valencia orange groves on Swingle citrumelo rootstock for three seasons has maintained soil pH below 6.5 on the flatwoods and 6.0 on ridge. Over the last three seasons of survey, root density as an index of root heath has been sustained. Phytophthora populations remain below the damaging level in ridge groves and in flatwoods increase to damaging levels coincident with the fall root flush but drop back to non-damaging levels for remainder of the season. Compared to the 2013-14 season, yields in the ridge blocks have increased up to 4% and on the flatwoods have increased up to 22%.Growers using acidification treatments with sulfuric and/or N-phuric acid for the last 3 seasons report an average cost of $60 per acre. This cost will analyzed in relation to yield response to provide a cost benefit of acidification
Non-Technical Summary: Managements have been implemented to reduce soil, nutrient and water stress, and Phytophthora root rot. They include frequent irrigation cycles, fertigation and acidification of irrigation water and soil to reduce rhizosphere pH, and fungicides. Root density of trees under these practices fluctuates seasonally and annually but has not declined over the past 3 years. Trees managed with soil acidification and fertigation have steadily recovered in health and yield.
http://www.icc2016.com/images/icc2016/downloads/Abstract_Book_ICC_2016.pdf
- Author: Ben Faber
This is a great website to view plant nutrient symptoms by plant or by nutrient. It is of ornamental plants, but hey, once you can see it on one plant you will something very similar on avocado, lemon, apple, almond, lychee or whatever alphabetic fruit you work with. Kudos to University of Florida. Of course once you see what iron deficiency looks like, you'll be able to identify it on most other plants:
http://hort.ufl.edu/database/nutdef/index.shtml
Below is Boron toxicity from a grower who got too excited about correcting B deficiency,
- Author: Ben Faber
I just raked up all the leaves under the avocado and it looks so nice. PUT THEM RIGHT BACK. The avocado is shallow-rooted and really depends on the natural leaf mulch to protect its roots. In fact the roots will actually colonize the rotted leaves as if it were soil. This mulch is also a first line of defense against root rot. The decomposing leaves create a hostile environment to the microorganism that causes the disease. The mulch also helps to reduce evaporative loss of water and therefore reduces water needs. Avocado growers will actually spread mulch in cases where trees are too young to produce adequate leaf drop for mulch or in windy areas where mulch has blown away. The key to remember is that the mulch should be kept at least six inches away from the trunk to avoid collar rot which can be causes by keeping a moist mulch against the trunk.